Psilotum nudum, commonly known as the whisk fern, is a fernlike plant found in tropical and subtropical regions around the world. It lacks roots and has greatly reduced leaves, earning it the Latin name Psilotum nudum, which means “bare naked”. It is often found growing epiphytically in tropical areas, and in rock crevices in more temperate areas. In Japan, it was once cultivated in gardens as an ornamental plant, and in Hawaii it was used medicinally and for making traditional leis. It was also used as a small broom, made by tying a handful of its branches together. It may also be a good source of antimicrobial chemicals.

Synonyms Top

Scientific name	Authority	First published in
Tristeca nuda	(L.) P.Beauv. ex J.St.-Hil.	Expos. Fam. Nat. 1: 39 1805
Psilotum triquetrum	Sw.	J. Bot. (Schrader) 1800(2): 109 (1801)
Psilotum floridanum	Michx.	Fl. Bor.-Amer. 2: 281 (1803)
Lycopodium nudum	L.	Sp. Pl. : 1100 (1753)
Psilotum triquetrum var. gracile	Grev. & Hook.	Bot. Misc. 2: 362 (1831)
Bernhardia antillarum	Müll.Berol.	Bot. Zeitung (Berlin) 14: 234 (1856)
Bernhardia deppeana	Müll.Berol.	Bot. Zeitung (Berlin) 14: 235 (1856)
Psilotum domingense	Gand.	Bull. Soc. Bot. France 66: 306 (1919)
Bernhardia novae-hollandiae	K.Mull.	Botanische Zeitung 14 1856 [28 Mar 1856]
Bernhardia oahuensis	Müll.Berol.	Bot. Zeitung (Berlin) 14: 235 (1856)
Psilotum triquetrum f. insulare	Brade	Bradea 1(1): 9, tab. 5 (1969)
Psilotum flabellatum	Gand.	Bulletin de la Soci?t? Botanique de France 66 66. 1919
Psilotum nudum var. fallicinum	(Domin) Domin	Biblioth. Bot. 22(89): (1928)
Psilotum triquetrum var. fallacinum	Domin	Repertorium Specierum Novarum Regni Vegetabilis 9 9. 1911 [15 Aug. 1911]
Psilotum nudum var. molesworthii	Iranzo, Prada & Salvo	Fl. Pterid. Andalucia 117. 1982

Common names Top

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Filter by language:

Language	Common/alternative name
English	upright whiskfern
English	whisk fern
English	forest moa
English	cocks crow
Spanish	canuto (cádiz)
Afrikaans	stoffervaring
Czech	prutovka nahá
Finnish	haarusanikkainen
Hawaiian	moa
Hawaiian	moa nehele
Hawaiian	pipi
Hawaiian	ʻoʻō moa
Hawaiian	oo moa
Japanese	マツバラン科
Japanese	マツバラン
Korean	솔잎란
Tamil	சைலோட்டம் நூடம்
Tonga	limufonua
Chinese	松叶蕨
Chinese	松葉蕨
Chinese	石刷把
Chinese	松叶兰

Subspecies (abbr. subsp./ssp.) Top

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Varieties (abbr. var.) Top

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Subvarieties (abbr. subvar.) Top

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Forms (abbr. f.) Top

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Germination/Propagation Top

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No germination or propagation data was added yet.

Distribution (via POWO/KEW) Top

Legend for the distribution data:

- Doubtful data

- Extinct

- Introduced

- Native

Africa click to expand
- East Tropical Africa
  - Kenya
  - Tanzania
  - Uganda
- Macaronesia
  - Canary Islands
  - Cape Verde
- South Tropical Africa
  - Angola
  - Mozambique
  - Zambia
  - Zimbabwe
- Southern Africa
  - Cape Provinces
  - Free State
  - Kwazulu-Natal
  - Lesotho
  - Northern Provinces
  - Swaziland
- West Tropical Africa
  - Ghana
  - Liberia
  - Nigeria
  - Senegal
- West-central Tropical Africa
  - Equatorial Guinea
  - Gabon
  - Gulf Of Guinea Islands
  - Zaïre
- Western Indian Ocean
  - Chagos Archipelago
  - Comoros
  - Madagascar
  - Mauritius
  - Rodrigues
  - Réunion
  - Seychelles

Asia-temperate click to expand
- Arabian Peninsula
  - Saudi Arabia
  - Yemen
- China
  - China North-central
  - China South-central
  - China Southeast
- Eastern Asia
  - Japan
  - Kazan-retto
  - Korea
  - Nansei-shoto
  - Ogasawara-Shoto
  - Taiwan

Asia-tropical click to expand
- Indian Subcontinent
  - Assam
  - Bangladesh
  - East Himalaya
  - India
  - Nepal
  - Pakistan
  - Sri Lanka
  - West Himalaya
- Indo-China
  - Andaman Islands
  - Laos
  - Myanmar
  - Nicobar Nicobar
  - South China Sea
  - Thailand
  - Vietnam
- Malesia
  - Christmas Island
  - Lesser Sunda Islands
  - Malaya
  - Maluku
  - Philippines
  - Sulawesi
- Papuasia
  - New Guinea
  - Solomon Islands

Australasia click to expand
- Australia
  - New South Wales
  - Norfolk Island
  - Northern Territory
  - Queensland
  - Victoria
  - Western Australia
- New Zealand
  - Kermadec Islands
  - New Zealand North

Europe click to expand
- Southwestern Europe
  - Spain

Northern America click to expand
- Mexico
  - Mexican Pacific Islands
  - Mexico Central
  - Mexico Gulf
  - Mexico Northeast
  - Mexico Northwest
  - Mexico Southeast
  - Mexico Southwest
- South-central U.S.A.
  - Texas
- Southeastern U.S.A.
  - Alabama
  - Arkansas
  - Florida
  - Georgia
  - Louisiana
  - Mississippi
  - North Carolina
  - South Carolina
- Southwestern U.S.A.
  - Arizona

Pacific click to expand
- North-central Pacific
  - Hawaii
- Northwestern Pacific
  - Caroline Islands
  - Marianas
  - Marshall Islands
- South-central Pacific
  - Cook Islands
  - Line Islands
  - Marquesas
  - Pitcairn Islands
  - Society Islands
  - Tuamotu
  - Tubuai Islands
- Southwestern Pacific
  - Fiji
  - Gilbert Islands
  - Nauru
  - New Caledonia
  - Niue
  - Samoa
  - Santa Cruz Island
  - Tokelau-manihiki
  - Tonga
  - Tuvalu
  - Vanuatu
  - Wallis-Futuna Islands

Southern America click to expand
- Brazil
  - Brazil North
  - Brazil Northeast
  - Brazil South
  - Brazil Southeast
  - Brazil West-central
- Caribbean
  - Bahamas
  - Bermuda
  - Cayman Islands
  - Cuba
  - Dominican Republic
  - Haiti
  - Jamaica
  - Leeward Islands
  - Puerto Rico
  - Venezuelan Antilles
  - Windward Islands
- Central America
  - Belize
  - Central American Pacific
  - Costa Rica
  - El Salvador
  - Guatemala
  - Honduras
  - Nicaragua
  - Panamá
- Northern South America
  - Venezuela
- Southern South America
  - Argentina Northeast
  - Argentina Northwest
  - Paraguay
  - Uruguay
- Western South America
  - Bolivia
  - Colombia
  - Ecuador
  - Galápagos
  - Peru

Links to other databases Top

Suggest others/fix!

Database	ID/link to page
World Flora Online	wfo-0001109597
Florida Plant Atlas	2108
Flora of Alabama	5252
USDA Plants	PSNU
Tropicos	26602814
INPN	448067
KEW	urn:lsid:ipni.org:names:212103-2
The Plant List	tro-26602814
Open Tree Of Life	144810
Observations.org	135994
NCBI Taxonomy	3240
Nature Serve	2.129404
IUCN Red List	18821051
IPNI	212103-2
iNaturalist	167517
GBIF	2688176
Freebase	/m/04f75dm
EPPO	QSINU
EOL	599703
Calflora (Californian flora)	10674
USDA GRIN	30219
Wikipedia	Psilotum_nudum
CMAUP	NPO23924

Genomes (via NCBI) Top

No reference genome is available on NCBI yet. We are constantly monitoring for new data.

Scientific Literature Top

Below are displayed the latest 15 articles published in PMC (PubMed Central®) and other sources (DOI number only)!
If you wish to see all the related articles click here.

Title

Authors

Publication

Released

IDs

Assembly and comparative analysis of the complete multichromosomal mitochondrial genome of Cymbidium ensifolium, an orchid of high economic and ornamental value

Shen B, Shen A, Liu L, Tan Y, Li S, Tan Z

BMC Plant Biol

09-Apr-2024

PMCID:	PMC11003039
doi:	10.1186/s12870-024-04962-4
PMID:	38594641

Telomere Length Variation in Model Bryophytes

Valeeva LR, Sannikova AV, Shafigullina NR, Abdulkina LR, Sharipova MR, Shakirov EV

Plants (Basel)

28-Jan-2024

PMCID:	PMC10856949
doi:	10.3390/plants13030387
PMID:	38337920

PMAT: an efficient plant mitogenome assembly toolkit using low-coverage HiFi sequencing data

Bi C, Shen F, Han F, Qu Y, Hou J, Xu K, Xu LA, He W, Wu Z, Yin T

Hortic Res

26-Jan-2024

PMCID:	PMC10925850
doi:	10.1093/hr/uhae023
PMID:	38469379

Worldwide Population Genomics Reveal Long-Term Stability of the Mitochondrial Genome Architecture in a Keystone Marine Plant

Khachaturyan M, Reusch TB, Dagan T

Genome Biol Evol

14-Sep-2023

PMCID:	PMC10538256
doi:	10.1093/gbe/evad167
PMID:	37708410

Transcriptome-Based Study on the Phylogeny and Hybridization of Marattialean Ferns (Marattiaceae)

Zhao J, Zhou X, Fang S, Zhu Z, Li Y, Yu H, He Z

Plants (Basel)

07-Jun-2023

PMCID:	PMC10301251
doi:	10.3390/plants12122237
PMID:	37375862

Characterizing conflict and congruence of molecular evolution across organellar genome sequences for phylogenetics in land plants

Tyszka AS, Bretz EC, Robertson HM, Woodcock-Girard MD, Ramanauskas K, Larson DA, Stull GW, Walker JF

Front Plant Sci

30-Mar-2023

PMCID:	PMC10098128
doi:	10.3389/fpls.2023.1125107
PMID:	37063179

Rickettsial DNA and a trans-splicing rRNA group I intron in the unorthodox mitogenome of the fern Haplopteris ensiformis

Zumkeller S, Polsakiewicz M, Knoop V

Commun Biol

20-Mar-2023

PMCID:	PMC10027690
doi:	10.1038/s42003-023-04659-8
PMID:	36941328

Categorizing 161 plant (streptophyte) mitochondrial group II introns into 29 families of related paralogues finds only limited links between intron mobility and intron-borne maturases

Zumkeller S, Knoop V

BMC Ecol Evol

13-Mar-2023

PMCID:	PMC10012718
doi:	10.1186/s12862-023-02108-y
PMID:	36915058

Giant Fern Genomes Show Complex Evolution Patterns: A Comparative Analysis in Two Species of Tmesipteris (Psilotaceae)

Fernández P, Leitch IJ, Leitch AR, Hidalgo O, Christenhusz MJ, Pokorny L, Pellicer J

Int J Mol Sci

31-Jan-2023

PMCID:	PMC9916801
doi:	10.3390/ijms24032708
PMID:	36769031

Light, rather than circadian rhythm, regulates gas exchange in ferns and lycophytes

Aros-Mualin D, Guadagno CR, Silvestro D, Kessler M

Plant Physiol

24-Jan-2023

PMCID:	PMC10022864
doi:	10.1093/plphys/kiad036
PMID:	36691320

Tissue-Specific Profiling of Biflavonoids in Ginkgo (Ginkgo biloba L.)

Kovač Tomas M, Jurčević I, Šamec D

Plants (Basel)

28-Dec-2022

PMCID:	PMC9824678
doi:	10.3390/plants12010147
PMID:	36616276

Stand structure adjustment influences the biomass allocation in naturally generated Pinus massoniana seedlings through environmental factors

He W, Wang Y, Wang X, Wen X, Li T, Ye M, Chen G, Zhao K, Hou G, Li X, Fan C

Front Plant Sci

28-Oct-2022

PMCID:	PMC9650532
doi:	10.3389/fpls.2022.997795
PMID:	36388556

The Complete Mitochondrial Genome of Ophioglossum vulgatum L. Is with Highly Repetitive Sequences: Intergenomic Fragment Transfer and Phylogenetic Analysis

Hao J, Liang Y, Su Y, Wang T

Genes (Basel)

21-Jul-2022

PMCID:	PMC9316493
doi:	10.3390/genes13071287
PMID:	35886070

Phylotranscriptomics Illuminates the Placement of Whole Genome Duplications and Gene Retention in Ferns

Pelosi JA, Kim EH, Barbazuk WB, Sessa EB

Front Plant Sci

14-Jul-2022

PMCID:	PMC9330400
doi:	10.3389/fpls.2022.882441
PMID:	35909764

The first step into phenolic metabolism in the hornwort Anthoceros agrestis: molecular and biochemical characterization of two phenylalanine ammonia-lyase isoforms

Pezeshki S, Warmbier I, Busch T, Bauerbach E, Szövenyi P, Petersen M

Planta

07-Jul-2022

PMCID:	PMC9262799
doi:	10.1007/s00425-022-03944-w
PMID:	35796843

Phytochemical Profile Top

Add a new one!

Below are displayed the proven (via scientific papers) natural compounds!
You can also contribute to this by clicking here.

Name	PubChem ID	Canonical SMILES	MW	Found in	Proof
> Benzenoids / Benzene and substituted derivatives
1-Phenyl-5-heptene-1,3-diyne	5281154	Click to see CC=CC#CC#CC1=CC=CC=C1	166.22	unknown	via CMAUP database
Phenylethyl Alcohol	6054	Click to see C1=CC=C(C=C1)CCO	122.16	unknown	via CMAUP database
Phenylheptatriyne	77981	Click to see CC#CC#CC#CC1=CC=CC=C1	164.20	unknown	via CMAUP database
> Benzenoids / Benzene and substituted derivatives / Benzoic acids and derivatives / Hydroxybenzoic acid derivatives / Gallic acid and derivatives
Syringic acid	10742	Click to see COC1=CC(=CC(=C1O)OC)C(=O)O	198.17	unknown	via CMAUP database
> Benzenoids / Phenols / Methoxyphenols
Syringaldehyde	8655	Click to see COC1=CC(=CC(=C1O)OC)C=O	182.17	unknown	via CMAUP database
> Lignans, neolignans and related compounds / Furanoid lignans
Sesamin	72307	Click to see C1C2C(COC2C3=CC4=C(C=C3)OCO4)C(O1)C5=CC6=C(C=C5)OCO6	354.40	unknown	via CMAUP database
Yangambin	443028	Click to see COC1=CC(=CC(=C1OC)OC)C2C3COC(C3CO2)C4=CC(=C(C(=C4)OC)OC)OC	446.50	unknown	via CMAUP database
> Lipids and lipid-like molecules / Fatty Acyls / Fatty acid esters
2-Butoxyethyl oleate	6436064	Click to see CCCCCCCCC=CCCCCCCCC(=O)OCCOCCCC	382.60	unknown	via CMAUP database
> Lipids and lipid-like molecules / Fatty Acyls / Fatty acyl glycosides / Fatty acyl glycosides of mono- and disaccharides
(2R,3S,4S,5R,6R)-2-(hydroxymethyl)-6-[(2R)-1-hydroxytrideca-5,7,9,11-tetrayn-2-yl]oxyoxane-3,4,5-triol	54671425	Click to see CC#CC#CC#CC#CCCC(CO)OC1C(C(C(C(O1)CO)O)O)O	362.40	unknown	via CMAUP database
> Lipids and lipid-like molecules / Fatty Acyls / Fatty alcohol esters
(5E)-1-Phenyl-5-heptene-1,3-diyn-7-ol acetate	23274467	Click to see CC(=O)OCC=CC#CC#CC1=CC=CC=C1	224.25	unknown	via CMAUP database
7-Phenyl-2,4,6-heptatriyne-1-ol acetate	11644243	Click to see CC(=O)OCC#CC#CC#CC1=CC=CC=C1	222.24	unknown	via CMAUP database
> Lipids and lipid-like molecules / Fatty Acyls / Fatty alcohols
(R)-7-Phenyl-4,6-heptadiyne-1,2-diol	101808987	Click to see C1=CC=C(C=C1)C#CC#CCC(CO)O	200.23	unknown	via CMAUP database
(S)-7-Phenyl-4,6-heptadiyne-2-ol	101395939	Click to see CC(CC#CC#CC1=CC=CC=C1)O	184.23	unknown	via CMAUP database
7-Phenyl-2,4,6-heptatriyn-1-ol	3085176	Click to see C1=CC=C(C=C1)C#CC#CC#CCO	180.20	unknown	via CMAUP database
7-Phenylhept-2-en-4,6-diyn-1-ol	114662	Click to see C1=CC=C(C=C1)C#CC#CC=CCO	182.22	unknown	via CMAUP database
> Lipids and lipid-like molecules / Fatty Acyls / Lineolic acids and derivatives
2-Butoxyethyl linoleate	87553426	Click to see CCCCCC=CCC=CCCCCCCCC(=O)OCCOCCCC	380.60	unknown	via CMAUP database
2-Butoxyethyl linolenate	87552578	Click to see CCCCOCCOC(=O)CCCCCCCC=CCC=CCC=CCC	378.60	unknown	via CMAUP database
Ethyl Linoleate	5282184	Click to see CCCCCC=CCC=CCCCCCCCC(=O)OCC	308.50	unknown	via CMAUP database
Ethyl linolenate	5367460	Click to see CCC=CCC=CCC=CCCCCCCCC(=O)OCC	306.50	unknown	via CMAUP database
Linoleic Acid	5280450	Click to see CCCCCC=CCC=CCCCCCCCC(=O)O	280.40	unknown	via CMAUP database
Linolenic Acid	5280934	Click to see CCC=CCC=CCC=CCCCCCCCC(=O)O	278.40	unknown	via CMAUP database
Methyl linolenate	5319706	Click to see CCC=CCC=CCC=CCCCCCCCC(=O)OC	292.50	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Diterpenoids
Rrr-alpha-tocopherylquinone	46183941	Click to see CC1=C(C(=O)C(=C(C1=O)C)CCC(C)(CCCC(C)CCCC(C)CCCC(C)C)O)C	446.70	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Diterpenoids / Acyclic diterpenoids
(7S,11S)-3,7,11,15-Tetramethyl-2-hexadecene-1-ol	40467768	Click to see CC(C)CCCC(C)CCCC(C)CCCC(=CCO)C	296.50	unknown	via CMAUP database
(E,7S,11S)-3,7,11,15-tetramethylhexadec-2-enoic acid	42433537	Click to see CC(C)CCCC(C)CCCC(C)CCCC(=CC(=O)O)C	310.50	unknown	via CMAUP database
[(E,7R,11R)-3,7,11,15-tetramethylhexadec-2-enyl] heptanoate	101918983	Click to see CCCCCCC(=O)OCC=C(C)CCCC(C)CCCC(C)CCCC(C)C	408.70	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Monoterpenoids / Acyclic monoterpenoids
(-)-Citronellol	7793	Click to see CC(CCC=C(C)C)CCO	156.26	unknown	via CMAUP database
(R)-(+)-Citronellal	75427	Click to see CC(CCC=C(C)C)CC=O	154.25	unknown	via CMAUP database
D-Citronellol	101977	Click to see CC(CCC=C(C)C)CCO	156.26	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Monoterpenoids / Menthane monoterpenoids
(-)-Isopulegol	170833	Click to see CC1CCC(C(C1)O)C(=C)C	154.25	unknown	via CMAUP database
1alpha,3beta,4beta-p-Menthane-3,8-diol	22214620	Click to see CC1CCC(C(C1)O)C(C)(C)O	172.26	unknown	via CMAUP database
Menthoglycol	19100	Click to see CC1CCC(C(C1)O)C(C)(C)O	172.26	unknown	via CMAUP database
Neomenthoglycol	9920491	Click to see CC1CCC(C(C1)O)C(C)(C)O	172.26	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Quinone and hydroquinone lipids / Vitamin E compounds / Tocopherols
Alpha-Tocopherol	14985	Click to see CC1=C(C2=C(CCC(O2)(C)CCCC(C)CCCC(C)CCCC(C)C)C(=C1O)C)C	430.70	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Sesquiterpenoids
alpha-Cadinol	10398656	Click to see CC1=CC2C(CCC(C2CC1)(C)O)C(C)C	222.37	unknown	via CMAUP database
T-Muurolol	3084331	Click to see CC1=CC2C(CCC(C2CC1)(C)O)C(C)C	222.37	unknown	via CMAUP database
trans-Calamenene	6429022	Click to see CC1CCC(C2=C1C=CC(=C2)C)C(C)C	202.33	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Triterpenoids
Betulinic Acid	64971	Click to see CC(=C)C1CCC2(C1C3CCC4C5(CCC(C(C5CCC4(C3(CC2)C)C)(C)C)O)C)C(=O)O	456.70	unknown	via CMAUP database
Betulonic acid	122844	Click to see CC(=C)C1CCC2(C1C3CCC4C5(CCC(=O)C(C5CCC4(C3(CC2)C)C)(C)C)C)C(=O)O	454.70	unknown	via CMAUP database
Erythrodiol	101761	Click to see CC1(CCC2(CCC3(C(=CCC4C3(CCC5C4(CCC(C5(C)C)O)C)C)C2C1)C)CO)C	442.70	unknown	via CMAUP database
Morolic acid	489944	Click to see CC1(CCC2(CCC3(C(C2=C1)CCC4C3(CCC5C4(CCC(C5(C)C)O)C)C)C)C(=O)O)C	456.70	unknown	via CMAUP database
Squalene	638072	Click to see CC(=CCCC(=CCCC(=CCCC=C(C)CCC=C(C)CCC=C(C)C)C)C)C	410.70	unknown	via CMAUP database
Ursolic Acid	64945	Click to see CC1CCC2(CCC3(C(=CCC4C3(CCC5C4(CCC(C5(C)C)O)C)C)C2C1C)C)C(=O)O	456.70	unknown	via CMAUP database
> Lipids and lipid-like molecules / Steroids and steroid derivatives / Cycloartanols and derivatives
Cycloeucalenol	101690	Click to see CC1C2CCC3C4(CCC(C4(CCC35C2(C5)CCC1O)C)C(C)CCC(=C)C(C)C)C	426.70	unknown	via CMAUP database
> Lipids and lipid-like molecules / Steroids and steroid derivatives / Stigmastanes and derivatives
alpha-Spinasterin	5281331	Click to see CCC(C=CC(C)C1CCC2C1(CCC3C2=CCC4C3(CCC(C4)O)C)C)C(C)C	412.70	unknown	via CMAUP database
Beta-Sitosterol	222284	Click to see CCC(CCC(C)C1CCC2C1(CCC3C2CC=C4C3(CCC(C4)O)C)C)C(C)C	414.70	unknown	via CMAUP database
Schottenol	441837	Click to see CCC(CCC(C)C1CCC2C1(CCC3C2=CCC4C3(CCC(C4)O)C)C)C(C)C	414.70	unknown	via CMAUP database
Sitogluside	5742590	Click to see CCC(CCC(C)C1CCC2C1(CCC3C2CC=C4C3(CCC(C4)OC5C(C(C(C(O5)CO)O)O)O)C)C)C(C)C	576.80	unknown	via CMAUP database
Stigmast-4-en-3-one	5484202	Click to see CCC(CCC(C)C1CCC2C1(CCC3C2CCC4=CC(=O)CCC34C)C)C(C)C	412.70	unknown	via CMAUP database
Tetraacetyldaucosterol	12895763	Click to see CCC(CCC(C)C1CCC2C1(CCC3C2CC=C4C3(CCC(C4)OC5C(C(C(C(O5)COC(=O)C)OC(=O)C)OC(=O)C)OC(=O)C)C)C)C(C)C	745.00	unknown	via CMAUP database
> Organic acids and derivatives / Carboxylic acids and derivatives / Dicarboxylic acids and derivatives
Butanedioate;hydron	21952380	Click to see [H+].[H+].C(CC(=O)[O-])C(=O)[O-]	118.09	unknown	via CMAUP database
Glutaric acid	743	Click to see C(CC(=O)O)CC(=O)O	132.11	unknown	via CMAUP database
> Organic oxygen compounds / Organooxygen compounds / Alcohols and polyols / Cyclitols and derivatives / Quinic acids and derivatives
(1R,3R,4S,5R)-3,4-bis[3-(3,4-dihydroxyphenyl)prop-2-enoyloxy]-1,5-dihydroxycyclohexane-1-carboxylic acid	460890	Click to see C1C(C(C(CC1(C(=O)O)O)OC(=O)C=CC2=CC(=C(C=C2)O)O)OC(=O)C=CC3=CC(=C(C=C3)O)O)O	516.40	unknown	via CMAUP database
(1S,3R,4R,5R)-3,4-Bis[3-(3,4-dihydroxyphenyl)prop-2-enoyloxy]-1,5-dihydroxycyclohexane-1-carboxylic acid	153946	Click to see C1C(C(C(CC1(C(=O)O)O)OC(=O)C=CC2=CC(=C(C=C2)O)O)OC(=O)C=CC3=CC(=C(C=C3)O)O)O	516.40	unknown	via CMAUP database
3,4-Dicaffeoylquinic acid	6474309	Click to see C1C(C(C(CC1(C(=O)O)O)OC(=O)C=CC2=CC(=C(C=C2)O)O)OC(=O)C=CC3=CC(=C(C=C3)O)O)O	516.40	unknown	via CMAUP database
> Organic oxygen compounds / Organooxygen compounds / Alcohols and polyols / Cyclitols and derivatives / Shikimic acids and derivatves
Shikimic acid	8742	Click to see C1C(C(C(C=C1C(=O)O)O)O)O	174.15	unknown	via CMAUP database
> Organic oxygen compounds / Organooxygen compounds / Carbohydrates and carbohydrate conjugates / Glycosyl compounds / Phenolic glycosides
2H-Pyran-2-one, 6-(4-(beta-D-glucopyranosyloxy)phenyl)-5,6-dihydro-, (S)-	6452048	Click to see C1C=CC(=O)OC1C2=CC=C(C=C2)OC3C(C(C(C(O3)CO)O)O)O	352.30	unknown	via CMAUP database
4-[[2-O-Acetyl-6-O-[3-(4-hydroxyphenyl)-1-oxo-2-propenyl]-beta-D-glucopyranosyl]oxy]benzeneacrylic acid	15071014	Click to see CC(=O)OC1C(C(C(OC1OC2=CC=C(C=C2)C=CC(=O)O)COC(=O)C=CC3=CC=C(C=C3)O)O)O	514.50	unknown	via CMAUP database
4-[[6-O-[3-(4-Hydroxyphenyl)-1-oxo-2-propenyl]-beta-D-glucopyranosyl]oxy]benzeneacrylic acid	15071011	Click to see C1=CC(=CC=C1C=CC(=O)OCC2C(C(C(C(O2)OC3=CC=C(C=C3)C=CC(=O)O)O)O)O)O	472.40	unknown	via CMAUP database
> Organic oxygen compounds / Organooxygen compounds / Carbonyl compounds / Aryl ketones / Aryl alkyl ketones
2-Acetylthiophene	6920	Click to see CC(=O)C1=CC=CS1	126.18	unknown	via CMAUP database
> Organoheterocyclic compounds / Thiophenes / 2,5-disubstituted thiophenes
5-(Phenylethynyl)thiophene-2-methanol	2822242	Click to see C1=CC=C(C=C1)C#CC2=CC=C(S2)CO	214.28	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Aurone flavonoids
(Z)-7-[[beta-D-Glucopyranosyl]oxy]-3',4',6-trihydroxyaurone	15071008	Click to see C1=CC(=C(C=C1C=C2C(=O)C3=C(O2)C(=C(C=C3)O)OC4C(C(C(C(O4)CO)O)O)O)O)O	448.40	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Cinnamic acids and derivatives / Hydroxycinnamic acids and derivatives / Coumaric acids and derivatives
2-Propenoic acid, 3-(3,4-dihydroxyphenyl)-, ethyl ester	66883	Click to see CCOC(=O)C=CC1=CC(=C(C=C1)O)O	208.21	unknown	via CMAUP database
Ethyl caffeate	5317238	Click to see CCOC(=O)C=CC1=CC(=C(C=C1)O)O	208.21	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Coumarins and derivatives / Furanocoumarins / Psoralens
Psoralen	6199	Click to see C1=CC(=O)OC2=CC3=C(C=CO3)C=C21	186.16	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / Flavans / Flavanones / Flavanonols
Aromadendrin	122850	Click to see C1=CC(=CC=C1C2C(C(=O)C3=C(C=C(C=C3O2)O)O)O)O	288.25	unknown	via CMAUP database
Fustin	5317435	Click to see C1=CC(=C(C=C1C2C(C(=O)C3=C(O2)C=C(C=C3)O)O)O)O	288.25	unknown	via CMAUP database
Garbanzol	442410	Click to see C1=CC(=CC=C1C2C(C(=O)C3=C(O2)C=C(C=C3)O)O)O	272.25	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / Flavones
Apigenin	5280443	Click to see C1=CC(=CC=C1C2=CC(=O)C3=C(C=C(C=C3O2)O)O)O	270.24	unknown	via CMAUP database
Luteolin	5280445	Click to see C1=CC(=C(C=C1C2=CC(=O)C3=C(C=C(C=C3O2)O)O)O)O	286.24	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / Flavones / Flavonols
Rhamnocitrin	5320946	Click to see COC1=CC(=C2C(=C1)OC(=C(C2=O)O)C3=CC=C(C=C3)O)O	300.26	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / Flavonoid glycosides / Flavonoid O-glycosides
2,3-Dihydroxy-4-[(E)-3-(3,4-dihydroxyphenyl)acryloyl]phenyl 4-O,6-O-diacetyl-beta-D-glucopyranoside	10840030	Click to see CC(=O)OCC1C(C(C(C(O1)OC2=C(C(=C(C=C2)C(=O)C=CC3=CC(=C(C=C3)O)O)O)O)O)O)OC(=O)C	534.50	unknown	via CMAUP database
2',3,3',4-Tetrahydroxy-4'-(6-O-acetyl-beta-D-glucopyranosyloxy)-trans-chalcone	14213555	Click to see CC(=O)OCC1C(C(C(C(O1)OC2=C(C(=C(C=C2)C(=O)C=CC3=CC(=C(C=C3)O)O)O)O)O)O)O	492.40	unknown	via CMAUP database
3',4'-Di(beta-D-glucopyranosyloxy)-2',3,4-trihydroxychalcone	102149400	Click to see C1=CC(=C(C=C1C=CC(=O)C2=C(C(=C(C=C2)OC3C(C(C(C(O3)CO)O)O)O)OC4C(C(C(C(O4)CO)O)O)O)O)O)O	612.50	unknown	via CMAUP database
4'-(3-O,4-O,6-O-Triacetyl-beta-D-glucopyranosyloxy)-2',3,3',4-tetrahydroxychalcone	14861259	Click to see CC(=O)OCC1C(C(C(C(O1)OC2=C(C(=C(C=C2)C(=O)C=CC3=CC(=C(C=C3)O)O)O)O)O)OC(=O)C)OC(=O)C	576.50	unknown	via CMAUP database
4'-[(6-O-beta-D-Glucopyranosyl-beta-D-glucopyranosyl)oxy]-2',3,3',4-tetrahydroxychalcone	14213552	Click to see C1=CC(=C(C=C1C=CC(=O)C2=C(C(=C(C=C2)OC3C(C(C(C(O3)COC4C(C(C(C(O4)CO)O)O)O)O)O)O)O)O)O)O	612.50	unknown	via CMAUP database
chrysosplenosid-C	118856019	Click to see COC1=C(C=CC(=C1)C2=C(C(=O)C3=C(C(=C(C=C3O2)OC)OC4C(C(C(C(O4)CO)O)O)O)O)OC)O	522.50	unknown	via CMAUP database
Marein	6441269	Click to see C1=CC(=C(C=C1C=CC(=O)C2=C(C(=C(C=C2)OC3C(C(C(C(O3)CO)O)O)O)O)O)O)O	450.40	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / Flavonoid glycosides / Flavonoid O-glycosides / Aurone O-glycosides
(2Z)-2-(3,4-Dihydroxybenzylidene)-6-[(4-O,6-O-diacetyl-beta-D-glucopyranosyl)oxy]-7-hydroxybenzofuran-3(2H)-one	101669623	Click to see CC(=O)OCC1C(C(C(C(O1)OC2=C(C3=C(C=C2)C(=O)C(=CC4=CC(=C(C=C4)O)O)O3)O)O)O)OC(=O)C	532.40	unknown	via CMAUP database
(Z)-6-[[6-O-[3-(4-Hydroxyphenyl)-1-oxo-2-propenyl]-beta-D-glucopyranosyl]oxy]-3',4',7-trihydroxyaurone	15071009	Click to see C1=CC(=CC=C1C=CC(=O)OCC2C(C(C(C(O2)OC3=C(C4=C(C=C3)C(=O)C(=CC5=CC(=C(C=C5)O)O)O4)O)O)O)O)O	594.50	unknown	via CMAUP database
(Z)-6-[[6-O-Acetyl-beta-D-glucopyranosyl]oxy]-3',4',7-trihydroxyaurone	15071010	Click to see CC(=O)OCC1C(C(C(C(O1)OC2=C(C3=C(C=C2)C(=O)C(=CC4=CC(=C(C=C4)O)O)O3)O)O)O)O	490.40	unknown	via CMAUP database
2-[(alphaZ)-3,4-Dihydroxybenzylidene]-6-(2-O,4-O,6-O-triacetyl-beta-D-glucopyranosyloxy)-7-hydroxybenzofuran-3(2H)-one	10674637	Click to see CC(=O)OCC1C(C(C(C(O1)OC2=C(C3=C(C=C2)C(=O)C(=CC4=CC(=C(C=C4)O)O)O3)O)OC(=O)C)O)OC(=O)C	574.50	unknown	via CMAUP database
2-[(alphaZ)-3,4-Dihydroxybenzylidene]-6-(3-O,4-O,6-O-triacetyl-beta-D-glucopyranosyloxy)-7-hydroxybenzofuran-3(2H)-one	10603216	Click to see CC(=O)OCC1C(C(C(C(O1)OC2=C(C3=C(C=C2)C(=O)C(=CC4=CC(=C(C=C4)O)O)O3)O)O)OC(=O)C)OC(=O)C	574.50	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / Flavonoid glycosides / Flavonoid O-glycosides / Flavonoid-3-O-glycosides
Rutin	5280805	Click to see CC1C(C(C(C(O1)OCC2C(C(C(C(O2)OC3=C(OC4=CC(=CC(=C4C3=O)O)O)C5=CC(=C(C=C5)O)O)O)O)O)O)O)O	610.50	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / Flavonoid glycosides / Flavonoid O-glycosides / Flavonoid-7-O-glycosides
3,3'-Dimethoxy-4',5-dihydroxy-7-(beta-D-glucopyranosyloxy)flavone	13344657	Click to see COC1=C(C=CC(=C1)C2=C(C(=O)C3=C(C=C(C=C3O2)OC4C(C(C(C(O4)CO)O)O)O)O)OC)O	492.40	unknown	via CMAUP database
3,3'-Dimethoxy-4',5-dihydroxy-7-[(6-O-alpha-L-rhamnopyranosyl-beta-D-glucopyranosyl)oxy]flavone	102316697	Click to see CC1C(C(C(C(O1)OCC2C(C(C(C(O2)OC3=CC(=C4C(=C3)OC(=C(C4=O)OC)C5=CC(=C(C=C5)O)OC)O)O)O)O)O)O)O	638.60	unknown	via CMAUP database
5-hydroxy-2-(3-hydroxy-4-methoxy-phenyl)-3-methoxy-7-[(2S,3R,4S,5S,6R)-3,4,5-trihydroxy-6-[[(2R,3R,4R,5R,6S)-3,4,5-trihydroxy-6-methyl-tetrahydropyran-2-yl]oxymethyl]tetrahydropyran-2-yl]oxy-chromen-4-one	5495545	Click to see CC1C(C(C(C(O1)OCC2C(C(C(C(O2)OC3=CC(=C4C(=C3)OC(=C(C4=O)OC)C5=CC(=C(C=C5)OC)O)O)O)O)O)O)O)O	638.60	unknown	via CMAUP database
Axillaroside	101243893	Click to see COC1=C(C=C2C(=C1O)C(=O)C(=C(O2)C3=CC(=C(C=C3)O)O)OC)OC4C(C(C(C(O4)CO)O)O)O	508.40	unknown	via CMAUP database
Centaurein	5489090	Click to see COC1=C(C=C(C=C1)C2=C(C(=O)C3=C(C(=C(C=C3O2)OC4C(C(C(C(O4)CO)O)O)O)OC)O)OC)O	522.50	unknown	via CMAUP database
Jacein	11577257	Click to see COC1=C(C=CC(=C1)C2=C(C(=O)C3=C(C(=C(C=C3O2)OC4C(C(C(C(O4)CO)O)O)O)OC)O)OC)O	522.50	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / O-methylated flavonoids / 6-O-methylated flavonoids
Jaceidin	5464461	Click to see COC1=C(C=CC(=C1)C2=C(C(=O)C3=C(O2)C=C(C(=C3O)OC)O)OC)O	360.30	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / O-methylated flavonoids / 7-O-methylated flavonoids
(2R,3R)-3,4'-Dihydroxy-5,7-dimethoxyflavone	10403456	Click to see COC1=CC2=C(C(=C1)OC)C(=O)C(C(O2)C3=CC=C(C=C3)O)O	316.30	unknown	via CMAUP database
Aromadendrin 7-methyl ether	181132	Click to see COC1=CC(=C2C(=C1)OC(C(C2=O)O)C3=CC=C(C=C3)O)O	302.28	unknown	via CMAUP database
Eucalyptin	76573	Click to see CC1=C(C2=C(C(=C1OC)C)OC(=CC2=O)C3=CC=C(C=C3)OC)O	326.30	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Linear 1,3-diarylpropanoids / Chalcones and dihydrochalcones / 2-Hydroxychalcones
Okanin	5281294	Click to see C1=CC(=C(C=C1C=CC(=O)C2=C(C(=C(C=C2)O)O)O)O)O	288.25	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Tannins / Hydrolyzable tannins
Ellagic Acid	5281855	Click to see C1=C2C3=C(C(=C1O)O)OC(=O)C4=CC(=C(C(=C43)OC2=O)O)O	302.19	unknown	via CMAUP database

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Psilotum nudum

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