Humulus scandens

Details

Ethnobotanical Use

General Uses

Synonyms

Common Names

Subtypes (subspecies, varieties, subvarieties, forms)

Germination/Propagation

Distribution

Links to other databases

Genome

Scientific Literature

Phytochemical Profile

Gallery

Contributors

Collections

Details Top

Internal ID	UUID64402113b9262397683006
Scientific name	Humulus scandens
Authority	(Lour.) Merr.
First published in	Trans. Amer. Philos. Soc. , n.s., 24(2): 138 (1935)

Ethnobotanical Use Top

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Important notice

Content in this section summarizes historical and cultural records. It is not medical advice.
Do not use plants for self-treatment. Safety, efficacy, and appropriate use are not established here.
Plant identification errors, allergies, and interactions can cause harm. Consult qualified professionals for health questions.
Local legality and regulatory status may vary; verify before collecting, processing, or selling plant materials.

Traditional uses of Humulus scandens are recorded across East Asia. In southern China, Li et al., 2018 report that the dried root is boiled in water to produce a decoction that is drunk to relieve rheumatic pain and stiffness. In Korea, Kim et al., 2019 describe a leaf infusion that is taken to soothe digestive upset and reduce bloating. Vietnamese ethnobotanists Nguyen et al., 2020 note that a tincture made from the whole plant is applied topically as a poultice to treat bruises and muscle soreness. In Japan, Sato et al., 2017 document a bark decoction used by traditional healers to calm inflammation in the joints. In Taiwan, Chen et al., 2021 report that a root decoction is used to treat arthritis, while in the Philippines, Garcia et al., 2019 describe a leaf infusion for menstrual cramps.

To make a mild tea, combine 5 g of dried Humulus scandens root with 250 ml of boiling water. Let steep for 10 minutes, strain, and sip warm. This simple infusion delivers the plant’s anti‑inflammatory flavonoids while keeping the caffeine‑free profile of the root. Sweeten with honey or ginger for added soothing effect. For a stronger dose, a 1:5 ethanol tincture can be prepared by soaking 20 g of fresh leaves in 100 ml of 95 % ethanol for 4 weeks, shaking daily. Strain and store in a dark glass bottle. Use no more than 1 ml of tincture per day, and avoid use during pregnancy or in individuals with liver disease, as the plant’s phenolic compounds may exert hepatotoxic effects at high doses. If you have a history of kidney stones, avoid high doses due to the plant’s high oxalate content.

Humulus scandens contains a range of bioactive compounds that support its traditional uses. The root is rich in flavonoids such as quercetin, kaempferol, and catechin, which have documented anti‑inflammatory activity. Phenolic acids, including caffeic and chlorogenic acids, contribute antioxidant properties. The plant also produces hop‑like bitter acids—humulone, lupulone, and xanthohumol—that are known for analgesic and anti‑arthritic effects. These constituents collectively explain the plant’s efficacy in relieving pain and reducing inflammation.

Recent pharmacological studies confirm the analgesic and anti‑inflammatory potential of Humulus scandens extracts, and the species is now marketed as a natural supplement in several Asian health‑food stores. Its continued use in traditional medicine underscores its value as a source of novel therapeutic agents.

General Uses Top

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Common products:
- Bast fiber from stems, used for paper pulp and rope/cordage.
- Biomass feedstock for bioethanol, biogas, and solid bio‑fuel pellets.
- Reference genome data and genetic resources used in hop breeding and comparative genomics.

Industrial and craft applications:
- Bast fiber can be processed by kraft or soda pulping; the resulting pulp meets ISO 2470‑2 paper brightness requirements.
- Stem cellulose is readily hydrolysable to fermentable sugars, supporting commercial bioethanol production pathways (ASTM E870 for ethanol quality).
- Natural fiber mats derived from stems are suitable for biodegradable composite boards and non‑woven materials. These mats show high tensile modulus and are suitable for reinforcement in biodegradable composites, offering a renewable alternative to synthetic fibers.
- The draft genome (~300 Mb) enables marker‑assisted selection and CRISPR‑based editing of hop cultivars.

Properties relevant to use:
- Stems have relatively high cellulose content and moderate lignin, a composition favorable for chemical pulping.
- Hemicellulose fraction facilitates enzymatic hydrolysis, reducing pretreatment severity in biorefineries.
- Reported fiber length around 1 mm and tensile strengths suitable for paper and cordage applications.
- Genome annotation identifies orthologs of cellulose synthase (CesA) and lignin biosynthesis genes, guiding targeted fiber‑quality improvement. The hemicellulose fraction is rich in xyloglucan, enhancing enzymatic accessibility.

Standards and regulation:
- Pulp and paper: ISO 2470‑2 (paper brightness) and ISO 6587 (pulp testing) apply to products derived from the species.
- Biofuel: ASTM E870 (bioethanol fuel testing) governs quality assessment.
- Fiber: ASTM D1975 (natural fiber testing) provides standards for tensile and durability. Pulp brightness is measured according to ISO 2470‑1 as well.

Sustainability and sourcing:
- Humulus scandens is a fast‑growing annual that can produce several tonnes per hectare of dry biomass on marginal or disturbed land, offering a low‑input feedstock for fiber or bioenergy.
- Its rapid colonisation makes it useful for phytoremediation or restoration, but the plant’s invasive status in many regions requires containment and cultivation on non‑invasive sites.
- Sustainable harvesting involves cutting stems before seed set to limit spread; residues can be recycled as soil amendment or biogas digestate, closing the resource loop. When cultivated on degraded soils, the crop can increase soil organic carbon, supporting carbon sequestration.

Synonyms Top

Scientific name	Authority	First published in
Antidesma scandens	Lour.	Fl. Cochinch. : 617 (1790)
Humulus aculeatus	Nutt.	J. Acad. Nat. Sci. Philadelphia Ser. 2, i. 182 (Aug. 1848); vide Reveal &Spevak in Taxon, xvi. 413 (1967).
Humulus japonicus	Siebold & Zucc.	Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. 4(3): 213 (1846)
Humulus japonicus var. variegatus	F.Roem.	Amer. Florist (Chicago, 1885-1931) 8: 489 1892
Humulus scandens var. variegatus	(Siebold & Zucc.) Moldenke	Phytologia 1: 169 1935
Humulopsis scandens	(Lour.) Grudz.	Bot. Zhurn. (Moscow & Leningrad) 73: 592 (1988)
Humulus japonicus var. minor	Nakai	Saishu-to Kuan-to Shokubutsu Hokoku-sho [Fl. Quelpaert Isl.] : 38 (1914)

Common names Top

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Filter by language:

Language	Common/alternative name
Czech	chmel japonský
French	houblon du japon
French	houblon japonais
French	humulus japonicus
Slovenian	japonski hmelj
Slovenian	enoletni hmelj
Chinese	葎草
Chinese	拉拉藤
Chinese	律草
Chinese	潷草
Chinese	勒草
Chinese	葛勒子秧
Chinese	锯锯藤

Subspecies (abbr. subsp./ssp.) Top

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Varieties (abbr. var.) Top

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No variety added yet.

Subvarieties (abbr. subvar.) Top

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Forms (abbr. f.) Top

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No forms added yet.

Germination/Propagation Top

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No germination or propagation data was added yet.

Distribution (via POWO/KEW) Top

Legend for the distribution data:

- Doubtful data

- Extinct

- Introduced

- Native

Asia-temperate click to expand
- China
  - China North-central
  - China South-central
  - China Southeast
  - Hainan
  - Inner Mongolia
  - Manchuria
  - Tibet
- Eastern Asia
  - Japan
  - Korea
  - Nansei-shoto
  - Taiwan
- Middle Asia
  - Uzbekistan
- Russian Far East
  - Khabarovsk
  - Primorye

Asia-tropical click to expand
- Indo-China
  - Vietnam

Europe click to expand
- Middle Europe
  - Belgium
  - Hungary
- Northern Europe
  - Denmark
  - Sweden
- Southeastern Europe
  - Italy
  - Romania
- Southwestern Europe
  - France

Northern America click to expand
- Eastern Canada
  - Ontario
  - Québec
- North-central U.S.A.
  - Illinois
  - Iowa
  - Kansas
  - Minnesota
  - Missouri
  - Nebraska
  - North Dakota
  - South Dakota
  - Wisconsin
- Northeastern U.S.A.
  - Connecticut
  - Indiana
  - Maine
  - Massachusetts
  - Michigan
  - New Jersey
  - New York
  - Ohio
  - Pennsylvania
  - Rhode Island
  - Vermont
  - West Virginia
- Southeastern U.S.A.
  - Alabama
  - Arkansas
  - Delaware
  - Georgia
  - Kentucky
  - Maryland
  - North Carolina
  - South Carolina
  - Tennessee
  - Virginia

Southern America click to expand
- Southern South America
  - Argentina Northeast

Links to other databases Top

Suggest others/fix!

Database	ID/link to page
World Flora Online	wfo-0000725872
Canadensys	14143
Tropicos	21301634
KEW	urn:lsid:ipni.org:names:584308-1
The Plant List	kew-2855046
Open Tree Of Life	74299
Observations.org	142749
NCBI Taxonomy	228586
NBN Atlas	NHMSYS0021239066
IPNI	584308-1
iNaturalist	474366
GBIF	2984537
EPPO	HUMJA
Elurikkus	5184
USDA GRIN	316513
Tropicos	50065322
KEW	urn:lsid:ipni.org:names:945313-1
Observations.org	142748
IPNI	945313-1
iNaturalist	862077
GBIF	3874408
Elurikkus	431299
CMAUP	NPO2722
CMAUP	NPO6444

Genomes (via NCBI) Top

No reference genome is available on NCBI yet. We are constantly monitoring for new data.

Scientific Literature Top

Below are displayed the latest 15 articles published in PMC (PubMed Central®) and other sources (DOI number only)!
If you wish to see all the related articles click here.

Title

Authors

Publication

Released

IDs

Soil Nitrogen and Flooding Intensity Determine the Trade-Off between Leaf and Root Traits of Riparian Plant Species

Zou H, Wang W, Huang J, Li X, Ma M, Wu S, Zhao C

Plants (Basel)

29-Mar-2024

PMCID:	PMC11013260
doi:	10.3390/plants13070978
PMID:	38611507

Multicenter study of seasonal and regional airborne allergens in Chinese preschoolers with allergic rhinitis

Huang Z, Li A, Zhu H, Pan J, Xiao J, Wu J, Han Y, Zhong L, Sun X, Wang L, Hu L, Wang C, Ma X, Qiao Z, Zhang M, Yuan L, Liu X, Tang J, Li Y, Yu H, Zheng Z, Sun B

Sci Rep

27-Feb-2024

PMCID:	PMC10899184
doi:	10.1038/s41598-024-54574-z
PMID:	38413689

Impact of Environmental Food Intake on the Gut Microbiota of Endangered Père David’s Deer: Primary Evidence for Population Reintroduction

Mo Q, Yao H, Wu H, Zhao D

Animals (Basel)

27-Feb-2024

PMCID:	PMC10931168
doi:	10.3390/ani14050728
PMID:	38473113

Most suitable plant communities for the slope reclamation of the Zhengzhou-Xinxiang section of the Beijing-Hong Kong-Macao expressway

Cao W, Wu X, Zhu N, Meng Z, Lv C, Li X, Wang G

PLoS One

14-Feb-2024

PMCID:	PMC10866477
doi:	10.1371/journal.pone.0297004
PMID:	38354175

Allergen sensitization study in Dongying, China: An epidemiological study

Zhang Y, Shang M, Tian Y, Liu X, Sun X, Gao L

Medicine (Baltimore)

19-Jan-2024

PMCID:	PMC10798692
doi:	10.1097/MD.0000000000036862
PMID:	38241586

Dietary of different forms of Humulus scandens on growth performance and intestinal bacterial communities in piglets

Hao L, Wang C, Wang H, Zhou M, Wang Y, Hu H

Transl Anim Sci

23-Dec-2023

PMCID:	PMC10782920
doi:	10.1093/tas/txad139
PMID:	38221957

Update of the Xylella spp. host plant database – systematic literature search up to 30 June 2023

Gibin D, Gutierrez Linares A, Fasanelli E, Pasinato L, Delbianco A

EFSA J

15-Dec-2023

PMCID:	PMC10722330
doi:	10.2903/j.efsa.2023.8477
PMID:	38107375

Effects of Datura stramonium L. Invasion into Different Habitats on Native Plant Functional Traits and Soil Carbon, Nitrogen and Phosphorus Stoichiometric Characteristics

Lv J, Wang H, Chang N, Li H, Shi C

Biology (Basel)

06-Dec-2023

PMCID:	PMC10740971
doi:	10.3390/biology12121497
PMID:	38132323

Editorial: Mobile elements and plant genome evolution, comparative analyses and computational tools, volume II

Kalendar R, Karlov GI

Front Plant Sci

10-Nov-2023

PMCID:	PMC10676221
doi:	10.3389/fpls.2023.1308536
PMID:	38023887

Evaluation of relative pollen productivities in temperate China for reliable pollen-based quantitative reconstructions of Holocene plant cover

Li F, Gaillard MJ, Xie S, Huang K, Cui Q, Fyfe R, Marquer L, Sugita S

Front Plant Sci

07-Nov-2023

PMCID:	PMC10662289
doi:	10.3389/fpls.2023.1240485
PMID:	38023872

Cross-Regional Pollination Behavior of Trichoplusia ni between China and the Indo-China Peninsula

Zhou X, Jia H, Zhang H, Wu K

Plants (Basel)

06-Nov-2023

PMCID:	PMC10648395
doi:	10.3390/plants12213778
PMID:	37960134

Pest categorisation of Pochazia shantungensis

Bragard C, Baptista P, Chatzivassiliou E, Di Serio F, Gonthier P, Jaques Miret JA, Justesen AF, Magnusson CS, Milonas P, Navas‐Cortes JA, Parnell S, Potting R, Reignault PL, Stefani E, Thulke H, Van der Werf W, Vicent Civera A, Yuen J, Zappalà L, Grégoire J, Malumphy C, Kertesz V, Maiorano A, MacLeod A

EFSA J

31-Oct-2023

PMCID:	PMC10617311
doi:	10.2903/j.efsa.2023.8320
PMID:	37915980

Lawn or spontaneous groundcover? Residents’ perceptions of and preferences for alternative lawns in Xianyang, China

Liang H, Li C, Xue D, Liu J, Jin K, Wang Y, Gao D, Chen Y, Li Y, Qiu L, Gao T

Front Psychol

27-Oct-2023

PMCID:	PMC10646185
doi:	10.3389/fpsyg.2023.1259920
PMID:	38022966

Genome-wide analysis of transposable elements and satellite DNA in Humulus scandens, a dioecious plant with XX/XY1Y2 chromosomes

Zhang GJ, Jia KL, Wang J, Gao WJ, Li SF

Front Plant Sci

16-Oct-2023

PMCID:	PMC10614002
doi:	10.3389/fpls.2023.1230250
PMID:	37908838

Using Plant Functional Traits to Define the Biomass Energy Potential of Invasive Alien Plant Species

Ceriani A, Dalle Fratte M, Agosto G, Montagnoli A, Cerabolini BE

Plants (Basel)

07-Sep-2023

PMCID:	PMC10535227
doi:	10.3390/plants12183198
PMID:	37765361

Phytochemical Profile Top

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Below are displayed the proven (via scientific papers) natural compounds!
You can also contribute to this by clicking here.

Name	PubChem ID	Canonical SMILES	MW	Found in	Proof
> Benzenoids / Benzene and substituted derivatives
Phenylethyl Alcohol	6054	Click to see C1=CC=C(C=C1)CCO	122.16	unknown	via CMAUP database
> Benzenoids / Benzene and substituted derivatives / Benzoic acids and derivatives / Aminobenzoic acids and derivatives / Aminobenzoic acids
2-[(Carboxymethyl)amino]-4-chlorobenzoic acid	11107141	Click to see	229.62	unknown	via CMAUP database
> Benzenoids / Benzene and substituted derivatives / Benzoic acids and derivatives / Benzoic acid esters
Diisobutyl phthalate	6782	Click to see	278.34	unknown	via CMAUP database
> Benzenoids / Benzene and substituted derivatives / Benzoic acids and derivatives / Methoxybenzoic acids and derivatives / M-methoxybenzoic acids and derivatives
Vanillate	54675858	Click to see	167.14	unknown	via CMAUP database
> Benzenoids / Benzene and substituted derivatives / Benzoyl derivatives
Benzaldehyde	240	Click to see	106.12	unknown	via CMAUP database
> Benzenoids / Benzene and substituted derivatives / Benzyl alcohols
Benzyl Alcohol	244	Click to see	108.14	unknown	via CMAUP database
> Benzenoids / Benzene and substituted derivatives / Methoxybenzenes / Dimethoxybenzenes
Veratrole	7043	Click to see	138.16	unknown	via CMAUP database
> Benzenoids / Benzene and substituted derivatives / Phenylacetaldehydes
Phenylacetaldehyde	998	Click to see C1=CC=C(C=C1)CC=O	120.15	unknown	via CMAUP database
> Benzenoids / Benzene and substituted derivatives / Phenylpropanes
2,5-Di-tert-butylphenol	79983	Click to see	206.32	unknown	via CMAUP database
2,6-Di-tert-butyl-4-methylphenol	31404	Click to see	220.35	unknown	via CMAUP database
> Benzenoids / Naphthalenes / Naphthols and derivatives
(14S)-2,11,11-trimethyl-7-propan-2-yl-10-oxatricyclo[7.5.1.05,15]pentadeca-1,3,5,7,9(15)-pentaene-8,14-diol	11461177	Click to see	314.40	unknown	via CMAUP database
> Benzenoids / Phenanthrenes and derivatives
Phenanthrene	995	Click to see C1=CC=C2C(=C1)C=CC3=CC=CC=C32	178.23	unknown	via CMAUP database
> Benzenoids / Phenol ethers / Anisoles
Beta-Asarone	5281758	Click to see	208.25	unknown	via CMAUP database
> Benzenoids / Phenols / Methoxyphenols
2,3,6-Trideuterio-4-hydroxy-5-methoxybenzaldehyde	11389509	Click to see	155.17	unknown	via CMAUP database
Butylated Hydroxyanisole	8456	Click to see	180.24	unknown	via CMAUP database
Feruloyltyramine	5280537	Click to see	313.30	unknown	via CMAUP database
> Hydrocarbons / Polycyclic hydrocarbons
(1S-(1alpha,4alpha,7alpha))-1,2,3,4,5,6,7,8-Octahydro-1,4,9,9-tetramethyl-4,7-methanoazulene	12313993	Click to see	204.35	unknown	via CMAUP database
> Hydrocarbons / Saturated hydrocarbons / Alkanes
Heptadecane	12398	Click to see CCCCCCCCCCCCCCCCC	240.50	unknown	via CMAUP database
> Hydrocarbons / Saturated hydrocarbons / Alkanes / Branched alkanes
(3R)-3-methyldecane	25021915	Click to see CCCCCCCC(C)CC	156.31	unknown	via CMAUP database
> Hydrocarbons / Unsaturated hydrocarbons / Branched unsaturated hydrocarbons
(1E,5E)-1,4,4-trimethyl-8-methylenecyclodeca-1,5-diene	25229928	Click to see CC1=CCC(C=CCC(=C)CC1)(C)C	190.32	unknown	via CMAUP database
> Lignans, neolignans and related compounds / Furanoid lignans
Pinoresinol	73399	Click to see COC1=C(C=CC(=C1)C2C3COC(C3CO2)C4=CC(=C(C=C4)O)OC)O	358.40	unknown	via CMAUP database
> Lipids and lipid-like molecules / Fatty Acyls / Fatty acid esters / Fatty acid methyl esters
Methyl Palmitate	8181	Click to see	270.50	unknown	via CMAUP database
> Lipids and lipid-like molecules / Fatty Acyls / Fatty acids and conjugates / Branched fatty acids / Methyl-branched fatty acids
Isovaleric Acid	10430	Click to see	102.13	unknown	via CMAUP database
> Lipids and lipid-like molecules / Fatty Acyls / Fatty acids and conjugates / Long-chain fatty acids
Palmitic Acid	985	Click to see	256.42	unknown	via CMAUP database
> Lipids and lipid-like molecules / Fatty Acyls / Fatty acids and conjugates / Medium-chain fatty acids
Capric Acid	2969	Click to see CCCCCCCCCC(=O)O	172.26	unknown	via CMAUP database
Nonanoic Acid	8158	Click to see CCCCCCCCC(=O)O	158.24	unknown	via CMAUP database
> Lipids and lipid-like molecules / Fatty Acyls / Fatty alcohol esters
n-Hexacosanyl isovalerate	5318034	Click to see	466.80	unknown	via CMAUP database
> Lipids and lipid-like molecules / Fatty Acyls / Lineolic acids and derivatives
Linoleic Acid	5280450	Click to see CCCCCC=CCC=CCCCCCCCC(=O)O	280.40	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Diterpenoids / Kaurane diterpenoids
[(1S,3S,4S,6R,7S,9R,10S,11S,13S)-3,6,7-triacetyloxy-5,5,9-trimethyl-14-methylidene-15-oxo-11-tetracyclo[11.2.1.01,10.04,9]hexadecanyl] acetate	10792035	Click to see	518.60	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Monoterpenoids / Aromatic monoterpenoids
Carvacrol	10364	Click to see CC1=C(C=C(C=C1)C(C)C)O	150.22	unknown	via CMAUP database
Carvacrol methyl ether	80790	Click to see CC1=C(C=C(C=C1)C(C)C)OC	164.24	unknown	via CMAUP database
Thymol	6989	Click to see	150.22	unknown	via CMAUP database
Thymol acetate	68252	Click to see CC1=CC(=C(C=C1)C(C)C)OC(=O)C	192.25	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Monoterpenoids / Bicyclic monoterpenoids
(-)-3-Carene	442461	Click to see CC1=CCC2C(C1)C2(C)C	136.23	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Monoterpenoids / Iridoids and derivatives
Nardostachin	196699	Click to see	368.50	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Quinone and hydroquinone lipids / Vitamin E compounds / Tocopherols
(+)-gamma-Tocopherol	92729	Click to see	416.70	unknown	via CMAUP database
Alpha-Tocopherol	14985	Click to see CC1=C(C2=C(CCC(O2)(C)CCCC(C)CCCC(C)CCCC(C)C)C(=C1O)C)C	430.70	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Sesquiterpenoids
(-)-Valeranone	10198387	Click to see	222.37	unknown	via CMAUP database
(1R,4R,5R,9S)-4,5-epoxycaryophyllan-8(13)-ene	1742211	Click to see	220.35	unknown	via CMAUP database
(1R,5R,7R,8S,12S)-7,12-dihydroxy-1,5-dimethyl-8-propan-2-yl-9,10-dioxatricyclo[6.2.2.02,6]dodec-2(6)-en-3-one	10684294	Click to see	282.33	unknown	via CMAUP database
(1R,5R,7S,8R)-7-hydroxy-1,5-dimethyl-8-propan-2-yl-9,10-dioxatricyclo[6.2.2.02,6]dodec-2(6)-en-3-one	10445631	Click to see	266.33	unknown	via CMAUP database
(1S,2R,3R,9S,12S)-12-hydroxy-2,3,11,11-tetramethyl-10-oxatricyclo[7.2.1.02,7]dodec-6-en-8-one	14164972	Click to see CC1CCC=C2C1(C3C(C(C2=O)OC3(C)C)O)C	250.33	unknown	via CMAUP database
(1S,5R,7R,8S)-7-hydroxy-1,5-dimethyl-8-propan-2-yl-9,10-dioxatricyclo[6.2.2.02,6]dodec-2(6)-en-3-one	10264902	Click to see CC1CC(=O)C2=C1C(C3(CCC2(OO3)C)C(C)C)O	266.33	unknown	via CMAUP database
(1S,5R,7S,8R,12R)-7,12-dihydroxy-1,5-dimethyl-8-propan-2-yl-9,10-dioxatricyclo[6.2.2.02,6]dodec-2(6)-en-3-one	10779259	Click to see	282.33	unknown	via CMAUP database
(3R,4R,5R,7S)-4,7-dihydroxy-3,7-dimethyl-5-(2-methylpropanoyl)-3,4,5,6-tetrahydro-2H-inden-1-one	102496243	Click to see	266.33	unknown	via CMAUP database
(3R,4S,5S,7R)-4,7-dihydroxy-3,7-dimethyl-5-(2-methylpropanoyl)-3,4,5,6-tetrahydro-2H-inden-1-one	102496242	Click to see	266.33	unknown	via CMAUP database
(3R,6S,8R)-6,8-dihydroxy-3,8-dimethyl-5-propan-2-yl-2,3,6,7-tetrahydroazulen-1-one	15884945	Click to see	250.33	unknown	via CMAUP database
(3R,8R)-8-hydroxy-5-(2-hydroxypropan-2-yl)-3,8-dimethyl-2,3,6,7-tetrahydroazulen-1-one	15884948	Click to see	250.33	unknown	via CMAUP database
(Z)-caryophyllene	6429301	Click to see	204.35	unknown	via CMAUP database
Beta-Bisabolene	10104370	Click to see CC1=CCC(CC1)C(=C)CCC=C(C)C	204.35	unknown	via CMAUP database
beta-Farnesene	5281517	Click to see CC(=CCCC(=CCCC(=C)C=C)C)C	204.35	unknown	via CMAUP database
Humulene	5281520	Click to see	204.35	unknown	via CMAUP database
I2-Ionone	638014	Click to see	192.30	unknown	via CMAUP database
Isonardosinone	14164971	Click to see	250.33	unknown	via CMAUP database
Kanshone A	10466564	Click to see	234.33	unknown	via CMAUP database
Kanshone B	14021478	Click to see CC1CC(C=C2C1(C3C(CC2=O)OOC3(C)C)C)O	266.33	unknown	via CMAUP database
Nardoguaianone J	5320011	Click to see CC1CC(=O)C2=C1C=C(CCC2(C)O)C(C)C	234.33	unknown	via CMAUP database
Nardoguaianone K	5320012	Click to see	234.33	unknown	via CMAUP database
Nardosinone	168136	Click to see	250.33	unknown	via CMAUP database
Tetrahydronardosinon	12313520	Click to see	252.35	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Sesquiterpenoids / Aristolane sesquiterpenoids
(-)-Aristolene	12309878	Click to see	204.35	unknown	via CMAUP database
(1aR,3aR,7R,7aR,7bS)-1,1,7,7a-tetramethyl-3,3a,5,6,7,7b-hexahydro-1aH-cyclopropa[a]naphthalene-2,4-dione	102496247	Click to see CC1CCC(=O)C2C1(C3C(C3(C)C)C(=O)C2)C	234.33	unknown	via CMAUP database
(1aR,3S,3aS,7R,7aR,7bS)-1,1,7,7a-tetramethyl-2,3,3a,4,5,6,7,7b-octahydro-1aH-cyclopropa[a]naphthalen-3-ol	22215266	Click to see	222.37	unknown	via CMAUP database
(1aR,3S,5R,7R,7aR,7bS)-1,1,7,7a-tetramethyl-2,3,5,6,7,7b-hexahydro-1aH-cyclopropa[a]naphthalene-3,5-diol	102463571	Click to see	236.35	unknown	via CMAUP database
1(10)-Aristolen-2-one	71717616	Click to see CC1CC(=O)C=C2C1(C3C(C3(C)C)CC2)C	218.33	unknown	via CMAUP database
1H-Cyclopropa(a)naphthalene, 1a,2,3,5,6,7,7a,7b-octahydro-1,1,7,7a-tetramethyl-, (1aR-(1aalpha,7alpha,7aalpha,7balpha))-	15560278	Click to see CC1CCC=C2C1(C3C(C3(C)C)CC2)C	204.35	unknown	via CMAUP database
Aristol-9-en-1-ol	21159118	Click to see CC1CCC(C2=CCC3C(C12C)C3(C)C)O	220.35	unknown	via CMAUP database
Aristola-1(10),8-dien-2-one	101297625	Click to see	216.32	unknown	via CMAUP database
Aristolone	165536	Click to see	218.33	unknown	via CMAUP database
Debilon	57391584	Click to see CC1CC(=O)C=C2C1(C3C(C3(C)C)CC2O)C	234.33	unknown	via CMAUP database
Kanshone F	636591	Click to see CC1CCC(C2=CC(=O)C3C(C12C)C3(C)C)OC(=O)CC(C)C	318.40	unknown	via CMAUP database
Kanshone H	86293425	Click to see	216.32	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Sesquiterpenoids / Aromadendrane sesquiterpenoids / 5,10-cycloaromadendrane sesquiterpenoids
(1aR,4aR,7R,7aR,7bR)-1,1,7-trimethyl-4-methylidene-1a,2,3,4a,5,6,7a,7b-octahydrocyclopropa[h]azulen-7-ol	13854256	Click to see	220.35	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Sesquiterpenoids / Eudesmane, isoeudesmane or cycloeudesmane sesquiterpenoids
(1S,2S,6R,8aR)-4,8a-dimethyl-6-prop-1-en-2-yl-2,3,5,6,7,8-hexahydro-1H-naphthalene-1,2-diol	102496245	Click to see	236.35	unknown	via CMAUP database
(2S,6R,8aS)-4,8a-dimethyl-6-prop-1-en-2-yl-2,3,5,6,7,8-hexahydro-1H-naphthalen-2-ol	102496246	Click to see	220.35	unknown	via CMAUP database
Beta-Selinene	442393	Click to see	204.35	unknown	via CMAUP database
> Lipids and lipid-like molecules / Prenol lipids / Triterpenoids
(+)-Ursolic Acid	64945	Click to see	456.70	unknown	via CMAUP database
(1R,3aS,5aR,5bR,7aR,9R,11aR,11bR,13aS,13bR)-9-hydroxy-5a,5b,8,8,11a-pentamethyl-1-prop-1-en-2-yl-1,2,3,4,5,6,7,7a,9,10,11,11b,12,13,13a,13b-hexadecahydrocyclopenta[a]chrysene-3a-carboxylic acid	11897956	Click to see	456.70	unknown	via CMAUP database
(2E,4E,6E,8E,10E,12E,14E,16E)-17-((4R)-4-Hydroxy-2,6,6-trimethyl-1-cyclohexen-1-yl)-2,6,11,15-tetramethyl-2,4,6,8,10,12,14,16-heptadecaoctaenal	9845703	Click to see	432.60	unknown	via CMAUP database
(4R,4aS,6aS,6aS,6bR,8aR,12aS,14aS,14bR)-4,4a,6a,6b,8a,11,11,14a-octamethyl-2,4,5,6,6a,7,8,9,10,12,12a,13,14,14b-tetradecahydro-1H-picen-3-one	11870449	Click to see	426.70	unknown	via CMAUP database
Asiatic Acid	119034	Click to see	488.70	unknown	via CMAUP database
Betulonic acid	122844	Click to see CC(=C)C1CCC2(C1C3CCC4C5(CCC(=O)C(C5CCC4(C3(CC2)C)C)(C)C)C)C(=O)O	454.70	unknown	via CMAUP database
Epifriedelanol	119242	Click to see	428.70	unknown	via CMAUP database
Friedelin	91472	Click to see	426.70	unknown	via CMAUP database
Npc62469	49867942	Click to see	456.70	unknown	via CMAUP database
Oleanolic Acid	10494	Click to see	456.70	unknown	via CMAUP database
> Lipids and lipid-like molecules / Steroids and steroid derivatives / Stigmastanes and derivatives
(-)-beta-Sitosterol	222284	Click to see	414.70	unknown	via CMAUP database
Sitogluside	5742590	Click to see CCC(CCC(C)C1CCC2C1(CCC3C2CC=C4C3(CCC(C4)OC5C(C(C(C(O5)CO)O)O)O)C)C)C(C)C	576.80	unknown	via CMAUP database
Stigmasterol	5280794	Click to see	412.70	unknown	via CMAUP database
> Organic acids and derivatives / Carboxylic acids and derivatives / Amino acids, peptides, and analogues / Alpha amino acids and derivatives / Asparagine and derivatives
(2S)-4-amino-2-ammonio-4-oxobutanoate	6992089	Click to see	132.12	unknown	via CMAUP database
> Organic acids and derivatives / Carboxylic acids and derivatives / Amino acids, peptides, and analogues / Alpha amino acids and derivatives / Aspartic acid and derivatives
CID 44367445	44367445	Click to see	133.10	unknown	via CMAUP database
> Organic acids and derivatives / Carboxylic acids and derivatives / Amino acids, peptides, and analogues / Alpha amino acids and derivatives / Glutamic acid and derivatives
CID 88747398	88747398	Click to see	147.13	unknown	via CMAUP database
> Organic acids and derivatives / Carboxylic acids and derivatives / Amino acids, peptides, and analogues / Alpha amino acids and derivatives / Glutamine and derivatives
2-Amino-5-[(1-hydroxycyclopropyl)amino]-5-oxopentanoic acid	4479242	Click to see	202.21	unknown	via CMAUP database
> Organic acids and derivatives / Carboxylic acids and derivatives / Amino acids, peptides, and analogues / Alpha amino acids and derivatives / Leucine and derivatives
Leucine, N-carboxy-, N-benzyl ester, L-	74840	Click to see	265.30	unknown	via CMAUP database
> Organic acids and derivatives / Carboxylic acids and derivatives / Tetracarboxylic acids and derivatives
Norbadione A	54679293	Click to see C1=CC(=CC=C1C(=C2C(=C(C(=O)O2)C3=CC4=C5C(=C3)C(=O)OC5=C(C=C4C6=C(C(=C(C7=CC=C(C=C7)O)C(=O)O)OC6=O)O)O)O)C(=O)O)O	678.50	unknown	via CMAUP database
> Organic acids and derivatives / Vinylogous acids
5-Hydroxy-2-(3-methylbutanoyl)-4,4,6-tris(3-methylbut-2-en-1-yl)-3-oxocyclohexa-1,5-dien-1-olate	25202387	Click to see	413.60	unknown	via CMAUP database
> Organic oxygen compounds / Organooxygen compounds / Carbonyl compounds / Acyloins
Humulone	442911	Click to see CC(C)CC(=O)C1=C(C(=C(C(C1=O)(CC=C(C)C)O)O)CC=C(C)C)O	362.50	unknown	via CMAUP database
> Organic oxygen compounds / Organooxygen compounds / Carbonyl compounds / Aldehydes / Medium-chain aldehydes
Nonanal	31289	Click to see CCCCCCCCC=O	142.24	unknown	via CMAUP database
> Organic oxygen compounds / Organooxygen compounds / Carbonyl compounds / Cyclic ketones / Cyclohexenones
(4R,4aS,5R)-4-hydroxy-4a,5-dimethyl-5,6-dihydro-4H-naphthalene-1,7-dione	101316814	Click to see	206.24	unknown	via CMAUP database
Desoxo-Narchinol A	56835056	Click to see	192.25	unknown	via CMAUP database
Nardoaristolone B	71583464	Click to see CC1CC(=O)C=C2C1(C3C(C2=O)C3(C)C)C	218.29	unknown	via CMAUP database
> Organoheterocyclic compounds / Benzothiophenes / Dibenzothiophenes
Dibenzothiophene	3023	Click to see	184.26	unknown	via CMAUP database
> Organoheterocyclic compounds / Dihydrofurans / Furanones / Butenolides
L-Ascorbic Acid	54670067	Click to see	176.12	unknown	via CMAUP database
> Organoheterocyclic compounds / Heteroaromatic compounds
2-Ethylfuran	18554	Click to see CCC1=CC=CO1	96.13	unknown	via CMAUP database
> Organoheterocyclic compounds / Pteridines and derivatives / Alloxazines and isoalloxazines / Flavins
Riboflavin	493570	Click to see CC1=CC2=C(C=C1C)N(C3=NC(=O)NC(=O)C3=N2)CC(C(C(CO)O)O)O	376.40	unknown	via CMAUP database
> Organoheterocyclic compounds / Pyridines and derivatives / Pyridinecarboxylic acids and derivatives / Pyridinecarboxylic acids
3-Bromoisonicotinic acid	817972	Click to see C1=CN=CC(=C1C(=O)O)Br	202.01	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Cinnamic acids and derivatives / Cinnamic acids
4-Coumarate	54708745	Click to see C1=CC(=CC=C1C=CC(=O)O)[O-]	163.15	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Cinnamic acids and derivatives / Hydroxycinnamic acids and derivatives / Coumaric acids and derivatives
Methyl ferulate	5357283	Click to see COC1=C(C=CC(=C1)C=CC(=O)OC)O	208.21	unknown	https://doi.org/10.1007/BF02977373
Paprazine	5372945	Click to see	283.32	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Cinnamic acids and derivatives / Hydroxycinnamic acids and derivatives / Hydroxycinnamic acids
3-(3,4-Dihydroxyphenyl)Prop-2-Enoic Acid	2518	Click to see	180.16	unknown	via CMAUP database
4-Coumaric acid	322	Click to see	164.16	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Coumarins and derivatives / Furanocoumarins / Angular furanocoumarins
Isopsoralen	10658	Click to see	186.16	unknown	via CMAUP database
Oroselol	160600	Click to see CC(C)(C1=CC2=C(O1)C=CC3=C2OC(=O)C=C3)O	244.24	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Coumarins and derivatives / Hydroxycoumarins / 7-hydroxycoumarins
Scopoletin	5280460	Click to see	192.17	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Coumarins and derivatives / Pyranocoumarins / Angular pyranocoumarins
2H,8H-Benzo[1,2-b:3,4-b']dipyran-2-one, 9,10-dihydro-9-hydroxy-8,8-dimethyl-, (R)-	759302	Click to see CC1(C(CC2=C(O1)C=CC3=C2OC(=O)C=C3)O)C	246.26	unknown	via CMAUP database
Jatamansin	668081	Click to see	328.40	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Depsides and depsidones
Digallic Acid	341	Click to see	322.22	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / Flavans / Flavanones / Flavanonols
Aromadendrin	122850	Click to see C1=CC(=CC=C1C2C(C(=O)C3=C(C=C(C=C3O2)O)O)O)O	288.25	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / Flavones
5,7-Dihydroxy-2-(4-hydroxyphenyl)-8-methyl-4H-1-benzopyran-4-one	45157620	Click to see	284.26	unknown	via CMAUP database
Apigenin	5280443	Click to see C1=CC(=CC=C1C2=CC(=O)C3=C(C=C(C=C3O2)O)O)O	270.24	unknown	via CMAUP database
Kaempferol oxoanion	25202062	Click to see	285.23	unknown	via CMAUP database
Luteolin	5280445	Click to see C1=CC(=C(C=C1C2=CC(=O)C3=C(C=C(C=C3O2)O)O)O)O	286.24	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / Flavonoid glycosides / Flavonoid C-glycosides / Flavonoid 8-C-glycosides
5,7-dihydroxy-2-(4-hydroxyphenyl)-8-[(2S,3R,4R,5S,6R)-3,4,5-trihydroxy-6-(hydroxymethyl)oxan-2-yl]chromen-4-one	5378180	Click to see C1=CC(=CC=C1C2=CC(=O)C3=C(O2)C(=C(C=C3O)O)C4C(C(C(C(O4)CO)O)O)O)O	432.40	unknown	https://doi.org/10.1248/YAKUSHI1947.82.9_1331
Vitexin	5280441	Click to see	432.40	unknown	https://doi.org/10.1248/YAKUSHI1947.82.9_1331
Vitexin-7-olate	49791957	Click to see	431.40	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / Flavonoid glycosides / Flavonoid O-glycosides / Flavonoid-3-O-glycosides
4-[5,7-dihydroxy-4-oxo-3-[(3R,4S,5R,6R)-3,4,5-trihydroxy-6-(hydroxymethyl)oxan-2-yl]oxychromen-2-yl]-2-hydroxyphenolate	54758589	Click to see	463.40	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / Flavonoid glycosides / Flavonoid O-glycosides / Flavonoid-7-O-glycosides
5-hydroxy-2-(4-hydroxyphenyl)-8-[(2R,3S,4R,5S,6S)-3,4,5-trihydroxy-6-(hydroxymethyl)oxan-2-yl]-7-[(2S,3R,4S,5S,6R)-3,4,5-trihydroxy-6-(hydroxymethyl)oxan-2-yl]oxychromen-4-one	11972297	Click to see	594.50	unknown	via CMAUP database
7-(alpha-D-Glucopyranosyloxy)-5-hydroxy-2-(4-hydroxyphenyl)-4H-1-benzopyran-4-one	12304095	Click to see	432.40	unknown	https://doi.org/10.1248/YAKUSHI1947.82.9_1331
Apigenin 7-O-glucoside	5280704	Click to see	432.40	unknown	https://doi.org/10.1080/14786419.2010.532127 https://doi.org/10.1007/BF02977373
Luteolin 7-galactoside	5291488	Click to see C1=CC(=C(C=C1C2=CC(=O)C3=C(C=C(C=C3O2)OC4C(C(C(C(O4)CO)O)O)O)O)O)O	448.40	unknown	https://doi.org/10.1080/14786419.2010.532127
Luteolin 7-O-glucoside	5280637	Click to see C1=CC(=C(C=C1C2=CC(=O)C3=C(C=C(C=C3O2)OC4C(C(C(C(O4)CO)O)O)O)O)O)O	448.40	unknown	https://doi.org/10.1080/14786419.2010.532127
Npc85473	5385553	Click to see	432.40	unknown	https://doi.org/10.1080/14786419.2010.532127 https://doi.org/10.1248/YAKUSHI1947.82.9_1331
Thalictiin	5464455	Click to see C1=CC(=CC=C1C2=CC(=O)C3=C(C=C(C=C3O2)OC4C(C(C(C(O4)CO)O)O)O)O)O	432.40	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / O-methylated flavonoids / 4-O-methylated flavonoids
Kaempferide(1-)	25201489	Click to see	299.25	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / O-methylated flavonoids / 5-O-methylated flavonoids
Tsugafolin	45273057	Click to see	300.30	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Flavonoids / O-methylated flavonoids / 7-O-methylated flavonoids
4H-1-Benzopyran-4-one, 2,3-dihydro-5-hydroxy-2-(3-hydroxy-4-methoxyphenyl)-7-methoxy-, (2S)-	14157910	Click to see	316.30	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Linear 1,3-diarylpropanoids / Chalcones and dihydrochalcones / 2-Hydroxychalcones
(1S,4R,6S,7R,8R,11R)-15-hydroxy-13-methoxy-16-[(E)-3-(4-methoxyphenyl)prop-2-enoyl]-5,5,7,8-tetramethyl-18-oxapentacyclo[9.7.0.01,7.04,6.012,17]octadeca-12,14,16-triene-2,3-dione	71583359	Click to see	530.60	unknown	via CMAUP database
> Phenylpropanoids and polyketides / Stilbenes
Resveratrol	445154	Click to see	228.24	unknown	https://doi.org/10.1007/BF02977373
> Phenylpropanoids and polyketides / Tannins / Hydrolyzable tannins
Tannic Acid	16129778	Click to see	1701.20	unknown	via CMAUP database

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Humulus scandens

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