Genus Timonius in Family Rubiaceae
In botanical taxonomy, a genus (plural genera) is a rank used to group closely related species within a family. In the hierarchy, genus sits below family and above species.
Genera are defined by shared morphological, anatomical, and genetic characteristics (for example, features of flowers, fruits, seeds, or leaves) that indicate a close evolutionary relationship among the species they contain.
Each genus can include one or more species. Examples include Rosa (roses) and Solanum (nightshades, including tomato and eggplant).
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Genus Description
Suggest a correction!Timonius DC. is a genus of trees and shrubs in the coffee family Rubiaceae. The number of accepted species is variable among databases, but a recent working estimate places the genus at about 230 species, with most taxa centered in tropical Southeast Asia and the Pacific and a few in the Indian Ocean and northern Australia (POWO, 2024; WFO, 2024; Global Biodiversity Information Facility, 2024). Timonius DC. is the correct author citation.
The genus is characterized by opposite leaves with well-developed, persistent interpetiolar stipules that often form a conical cap (a calyptra) protecting the young shoot apex. Indumentum ranges from glabrous to densely sericeous or pubescent. Inflorescences are usually axillary, ranging from solitary flowers to small fascicles or reduced cymes; flowers are often unisexual, sometimes dioecious, with a tubular to funnel-shaped white or pale corolla that is typically five-lobed. The inferior ovary has a bilocular or rarely multilocular ovary with axile placentation, developing into a fleshy to more or less dry drupe with pyrenes that are often laterally flattened and sometimes keeled or winged. Pollinators are not well documented, and while some species produce fruits attractive to frugivorous birds or bats, specific dispersal syndromes remain tentative and require focused observation. Chromosome counts for the genus are poorly known, and a reliable base number is not available from the current literature.
Species diversity is highest in New Guinea and associated islands, with secondary centers in Malesia and the western Pacific; several taxa are narrow island endemics. Plants typically occur in lowland to lower montane rainforest, coastal forest, and secondary growth, from sea level to mid-elevations; a few extend into drier monsoon or limestone settings. Molecular phylogenetic work has placed Timonius within the greater Gardenieae complex, and a recent broad study of Pacific rubiaceous lineages resolved Timonius among several small, chiefly Pacific genera, supporting a focused Pacific radiation (Wagner et al., 2021). Traditional sectional or subgeneric treatments exist in older literature, but they do not map cleanly onto recent molecular results, and the infrageneric limits remain unsettled. No widespread synonymization has achieved consensus, and databases generally maintain Timonius as distinct (POWO, 2024; WFO, 2024).
Human relevance remains limited: most species are components of forest vegetation rather than cultivated plants, though a few are occasionally retained in agroforestry or appear in traditional building or craft uses in island communities. The genus is not a major timber or crop group and is not invasive.
Conservation concerns center on habitat loss and sea-level rise for island taxa, coupled with incomplete red-list assessments for most species and insufficient field data on population status (IUCN Red List, 2024). Future work—combining targeted phylogenomics with better geographic sampling and conservation assessments—will be essential to refine species limits and safeguard narrow endemics.
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Timonius abanii (Jun H.Chen)
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Timonius affinis (A.Gray)
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Timonius akunensis (S.P.Darwin)
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Timonius albus (Volkens)
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Timonius alejandroanus (J.G.Chavez, Meve & Liede)
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Timonius alius (S.P.Darwin)
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Timonius amboinicus (Boerl.)
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Timonius amungwiwanensis (S.P.Darwin)
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Timonius anodon (Boerl.)
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Timonius appendiculatus (Merr.)
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Timonius arboreus (Elmer)
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Timonius argenteus (Valeton)
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Timonius auriculatus (Merr.)
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Timonius avenis (Valeton)
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Timonius balansanus (Schltr.)
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Timonius beamanii (K.M.Wong & Jun H.Chen)
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Timonius belensis (Merr. & L.M.Perry)
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Timonius billitonensis (Valeton)
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Timonius bismarckensis (S.P.Darwin)
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Timonius bogoriensis (Valeton)
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Timonius borneensis (Valeton)
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Timonius bosaviensis (S.P.Darwin)
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Timonius bougainvillensis (Merr. & L.M.Perry)
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Timonius bracteatus (Merr. & L.M.Perry)
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Timonius bracteoides (S.P.Darwin)
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Timonius branderhorstii (Valeton)
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Timonius bullatus (K.M.Wong & Jun H.Chen)
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Timonius carii (S.P.Darwin)
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Timonius carstensensis (Wernham)
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Timonius caudatifolius (Elmer)
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Timonius caudatus (Valeton)
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Timonius celebicus (Koord.)
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Timonius clathratus (S.P.Darwin)
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Timonius clementis (Merr.)
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Timonius confertiflorus (Merr.)
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Timonius corneri (K.M.Wong)
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Timonius corymbosus (Valeton)
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Timonius creber (S.P.Darwin)
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Timonius cryptophlebus (S.Moore)
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Timonius cuneatus (Warb.)
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Timonius cyathicalyx (S.P.Darwin)
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Timonius decipiens (Merr. & L.M.Perry)
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Timonius densiflorus (Valeton)
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Timonius diffusus (Ridl.)
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Timonius dolichophyllus ((Merr. & L.M.Perry) S.P.Darwin)
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Timonius dumagat (J.G.Chavez)
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Timonius elegans (S.P.Darwin)
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Timonius enderianus (Warb.)
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Timonius epiphyticus (Elmer)
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Timonius eremiticus (J.G.Chavez & Banag)
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Timonius esherianus (W.W.Sm.)
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Timonius eximius (Schltr.)
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Timonius ferrugineus (Merr.)
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Timonius filipes (Wernham)
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Timonius finlaysonianus (Hook.f.)
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Timonius flavescens (Baker)
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Timonius fuscus (Walp.)
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Timonius gammillii (Elmer)
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Timonius glabrior ((Valeton) K.M.Wong)
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Timonius gracilipes (Merr.)
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Timonius grandifolius (Valeton)
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Timonius hentyi (S.P.Darwin)
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Timonius heptamerus (Wernham)
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Timonius hirsutus (Merr.)
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Timonius hydrangeifolius ((Miq.) Boerl.)
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Timonius imitans (Merr. & L.M.Perry)
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Timonius insularis (S.P.Darwin)
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Timonius involucratus (Merr.)
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Timonius jobiensis (Wernham)
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Timonius kaniensis (Valeton)
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Timonius kinabaluensis (Jun H.Chen)
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Timonius klossii (Wernham)
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Timonius koikokoensis (Wernham)
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Timonius koordersii (Valeton)
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Timonius korrensis (Kaneh.)
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Timonius kostermansii (S.P.Darwin)
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Timonius laevigatus (Valeton)
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Timonius lagenifer (S.P.Darwin)
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Timonius lamii (S.P.Darwin)
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Timonius lanceolatus (Merr.)
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Timonius lasianthoides (Valeton)
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Timonius latifolius (K.Schum. & Lauterb.)
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Timonius laugerioides (Wernham)
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Timonius laxus (King & Gamble)
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Timonius ledermannii (Valeton)
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Timonius leopoldii (Jun H.Chen & K.M.Wong)
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Timonius leptocalyx (S.P.Darwin)
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Timonius longiflorus (Merr.)
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Timonius longifolius (Valeton)
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Timonius longistipulus (Merr.)
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Timonius longitubus (Merr. & L.M.Perry)
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Timonius macrophyllus ((Teijsm. & Binn.) Valeton)
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Timonius matangensis (Valeton)
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Timonius melanophloeus (Merr. & L.M.Perry)
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Timonius meridionalis (S.P.Darwin)
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Timonius merokensis (Wernham)
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Timonius minahassae (Koord.)
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Timonius minutifolius (Valeton)
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Timonius modestus (Merr. & L.M.Perry)
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Timonius mollis (Valeton)
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Timonius moluccanus (S.P.Darwin)
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Timonius montanus (Ridl.)
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Timonius morobensis (S.P.Darwin)
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Timonius multinervis (Valeton)
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Timonius neobritannicus (S.P.Darwin)
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Timonius neocaledonicus (Schltr. & K.Krause)
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Timonius nitens (Merr. & L.M.Perry)
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Timonius nitidus (Fern.-Vill.)
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Timonius nolitangere (J.G.Chavez, Alejandro & Meve)
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Timonius novoguineensis (Warb.)
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Timonius nudiceps (Valeton)
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Timonius nymannii (Valeton)
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Timonius oblanceolatus (Valeton)
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Timonius oblongus (Valeton)
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Timonius obovatus (Elmer)
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Timonius octonervius (Merr.)
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Timonius oktediensis (S.P.Darwin)
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Timonius oligophlebius (Merr.)
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Timonius ophioliticus (Jun H.Chen)
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Timonius oreophilus (Ridl.)
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Timonius ovalis (Boerl.)
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Timonius ovalistipulus (Valeton)
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Timonius oxyphyllus (Gand.)
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Timonius pachyphyllus (Merr.)
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Timonius paiawensis (S.P.Darwin)
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Timonius palawanensis (Elmer)
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Timonius panayensis (Merr.)
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Timonius pannosus (Jun H.Chen)
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Timonius papuanus (Merr.)
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Timonius philippinensis (Merr.)
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Timonius platycarpus (Montrouz.)
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Timonius polyneurus (Valeton)
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Timonius ponapensis (Valeton)
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Timonius pseudaffinis (S.P.Darwin)
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Timonius pseudoarboreus (J.G.Chavez)
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Timonius pseudocapitatus ((Scheff.) F.Muell.)
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Timonius pubistipulus (S.P.Darwin)
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Timonius pulgarensis (Elmer)
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Timonius pullenii (S.P.Darwin)
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Timonius pulposus (C.T.White)
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Timonius quinqueflorus (Merr.)
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Timonius reflexus (S.P.Darwin)
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Timonius repertus (S.P.Darwin)
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Timonius ridleyi (Merr.)
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Timonius ridsdalei (J.G.Chavez)
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Timonius rigidus ((Miq.) F.Muell.)
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Timonius rivularis (Merr. & L.M.Perry)
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Timonius roemeri (Valeton)
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Timonius rosselensis (S.P.Darwin)
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Timonius rotundus (Merr.)
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Timonius rufescens ((Miq.) Boerl.)
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Timonius rufus (Valeton)
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Timonius salicifolius (Valeton)
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Timonius salsedoi (Fosberg & Sachet)
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Timonius samarensis (Merr.)
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Timonius scaber (Valeton)
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Timonius scabriflorus ((Valeton) Merr. & L.M.Perry)
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Timonius schefferi (Valeton)
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Timonius schumannii (Koord. ex Valeton)
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Timonius secundiflorus (S.P.Darwin)
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Timonius sessilis (Valeton)
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Timonius singularis ((F.Muell.) L.S.Sm.)
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Timonius solomonensis (Merr. & L.M.Perry)
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Timonius spesvitarum (J.G.Chavez)
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Timonius splendens (Boerl.)
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Timonius stenolobus (Jun H.Chen & K.M.Wong)
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Timonius stevendarwinii (J.G.Chavez)
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Timonius subauritus (Valeton)
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Timonius subavenis ((Valeton) S.P.Darwin)
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Timonius subcoriaceus (Valeton)
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Timonius subsessilis (Valeton)
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Timonius sulitii (Merr. & Quisumb. ex J.G.Chavez & Tandang)
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Timonius sylvestris (S.Moore)
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Timonius tambuyukonensis (Jun H.Chen)
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Timonius ternifolius (Fern.-Vill.)
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Timonius teysmannii (Valeton)
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Timonius timon ((Spreng.) Merr.)
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Timonius tomentosus ((Valeton) S.P.Darwin)
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Timonius trichanthus (Merr. & L.M.Perry)
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Timonius trichocaulon (Boerl.)
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Timonius trichocladus (Merr. & L.M.Perry)
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Timonius trichophorus (Merr.)
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Timonius uniflorus ((C.F.Gaertn.) Govaerts)
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Timonius urdanetensis (Elmer)
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Timonius vaccinioides (Wernham ex Ridl.)
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Timonius valetonii (Elmer)
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Timonius versteegii (Valeton)
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Timonius villosus (Valeton)
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Timonius virgatus (Merr. & L.M.Perry)
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Timonius wallichianus (Valeton)
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Timonius wollastonii (Wernham)
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Timonius wrayi (King & Gamble)
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Timonius xanthocarpus (Merr. & L.M.Perry)
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Timonius zuckianus (S.P.Darwin)