Genus Isoetes in Family Isoetaceae
In botanical taxonomy, a genus (plural genera) is a rank used to group closely related species within a family. In the hierarchy, genus sits below family and above species.
Genera are defined by shared morphological, anatomical, and genetic characteristics (for example, features of flowers, fruits, seeds, or leaves) that indicate a close evolutionary relationship among the species they contain.
Each genus can include one or more species. Examples include Rosa (roses) and Solanum (nightshades, including tomato and eggplant).
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Genus Description
Suggest a correction!Isoetes (family Isoetaceae) is a cosmopolitan genus of heterosporous lycophytes containing approximately 350–380 accepted species worldwide. Its members inhabit freshwater environments ranging from low‑land ponds to high‑altitude lakes on all continents except Antarctica, with the highest richness in temperate regions of North America, Eurasia, and the Southern Hemisphere. The type species is Isoetes lacustris (L.) L., designated by Linnaeus (POWO, 2024; WFO, 2024).
Plants are perennial, rosette‑forming herbs that arise from a compact corm and bear a dense tuft of unbranched, cylindrical leaves that may reach 30 cm. Each leaf bears a single basally situated sporangium hidden beneath a membranous indusium, and a small ligule occurs near the leaf base. The genus is heterosporous: megaspores are relatively large (0.5–1 mm) with a reticulate surface, while microspores are minute (
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Isoetes × altonharvillii (Musselman & R.D.Bray)
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Isoetes × angeloi (D.F.Brunt. & K.L.McIntosh)
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Isoetes × blondeaui (D.F.Brunt. & P.C.Sokoloff)
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Isoetes × brittonii (D.F.Brunt. & W.C.Taylor)
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Isoetes × brochonii (Motelay)
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Isoetes × eatonii (R.Dodge)
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Isoetes × fernaldii (D.F.Brunt. & P.C.Sokoloff)
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Isoetes × gopalkrishnae (S.K.Singh, P.K.Shukla & N.K.Dubey)
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Isoetes × harveyi (A.A.Eaton)
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Isoetes × herb-wagneri (W.C.Taylor)
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Isoetes × hickeyi (W.C.Taylor & Luebke)
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Isoetes × jermyi (D.F.Brunt. & Rumsey)
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Isoetes × kareniae (D.F.Brunt. & P.C.Sokoloff)
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Isoetes × marensis (D.M.Britton & D.F.Brunt.)
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Isoetes × michinokuana (M.Takamiya, Mits.Watan. & K.Ono)
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Isoetes × novae\-angliae (D.F.Brunt. & D.M.Britton)
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Isoetes × paratunica (D.F.Brunt., Mochalova & A.A.Bobrov)
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Isoetes × pseudotruncata (Britton & D.F.Brunt.)
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Isoetes × robusta ((Engelm.) D.F.Brunt.)
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Isoetes abyssinica (Chiov.)
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Isoetes aemulans (J.P.Roux)
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Isoetes aequinoctialis (Welw. ex A.Br.)
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Isoetes afloramientorum (D.F.Brunt. & J.B.S.Pereira)
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Isoetes alcalophila (S.Halloy)
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Isoetes alpina (Kirk)
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Isoetes alstonii (C.F.Reed & Verdc.)
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Isoetes amazonica (A.Braun)
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Isoetes anamariae (J.B.S.Pereira)
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Isoetes anatolica (Prada & Rolleri)
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Isoetes andicola ((Amstutz) L.D.Gómez)
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Isoetes andina (Spruce ex Hook.)
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Isoetes appalachiana (D.F.Brunt. & D.M.Britton)
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Isoetes araucaniana (Macluf & Hickey)
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Isoetes asiatica ((Makino) Makino)
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Isoetes attenuata (C.R.Marsden & Chinnock)
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Isoetes australis (S.Williams)
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Isoetes azorica (Durieu)
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Isoetes baculata (Hickey & H.P.Fuchs)
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Isoetes baodongii (Y.F.Gu, Y.H.Yan & Yi Jun Lu)
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Isoetes biafrana (Alston)
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Isoetes bilaspurensis (Panigrahi)
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Isoetes bischlerae (H.P.Fuchs)
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Isoetes bolanderi (Engelm.)
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Isoetes boliviensis (U.Weber)
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Isoetes boomii (Luebke)
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Isoetes boryana (Durieu)
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Isoetes boyacensis (H.P.Fuchs)
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Isoetes bradei (Herter)
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Isoetes brasiliensis (H.P.Fuchs)
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Isoetes brevicula (E.R.L.Johnson)
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Isoetes bruntonii (Knepper & Musselman)
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Isoetes butleri (Engelm.)
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Isoetes candelariensis (C.C.Macluf & G.J.Marquez)
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Isoetes cangae (J.B.S.Pereira, Salino & Stützel)
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Isoetes caparaoensis (J.B.S.Pereira)
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Isoetes capensis (A.V.Duthie)
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Isoetes carltaylori (Musselman)
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Isoetes caroli (E.R.L.Johnson)
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Isoetes caroliniana ((A.A.Eaton) Luebke)
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Isoetes changleensis (Yu C.Chen & Xing Liu)
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Isoetes changxingensis (Y.F.Gu & J.H.Shen)
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Isoetes chubutiana (Hickey, Macluf & W.C.Taylor)
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Isoetes cipoensis (J.B.S.Pereira)
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Isoetes coreana (Y.H.Chung & H.K.Choi)
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Isoetes coromandelina (L.f.)
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Isoetes creussensis (Lazare & S.Riba)
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Isoetes cristata (C.R.Marsden & Chinnock)
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Isoetes cubana (Engelm.)
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Isoetes delalandei (J.Lloyd)
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Isoetes delilei ((Bory) Rothm.)
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Isoetes dispora (Hickey)
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Isoetes dixitii (Shende)
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Isoetes dodgei (A.A.Eaton)
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Isoetes drummondii (A.Braun)
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Isoetes dubsii (J.B.S.Pereira)
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Isoetes duriei (Bory)
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Isoetes eatonii (R.Dodge)
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Isoetes echinospora (Durieu)
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Isoetes echtuckeri (D.F.Brunt. & D.M.Britton)
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Isoetes ecuadoriensis (Aspl.)
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Isoetes ekmanii (U.Weber)
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Isoetes elatior (A.Braun)
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Isoetes eludens (J.P.Roux, Hopper & Rhian J.Sm.)
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Isoetes engelmannii (A.Braun)
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Isoetes escondidensis (S.Halloy)
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Isoetes eshbaughii (Hickey)
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Isoetes fengii (Y.F.Gu & Y.H.Yan)
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Isoetes flaccida (A.Braun)
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Isoetes fluitans (M.I.Romero)
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Isoetes foveolata (A.A.Eaton in R.Dodge)
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Isoetes fuliginosa (R.L.Small & Hickey)
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Isoetes fuscomarginata (H.P.Fuchs)
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Isoetes gardneriana (Kunze)
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Isoetes georgiana (Luebke)
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Isoetes giessii (Launert)
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Isoetes gigantea (U.Weber)
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Isoetes graniticola (D.F.Brunt.)
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Isoetes gunnii (A.Braun)
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Isoetes gymnocarpa ((Gennari) A.Braun)
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Isoetes habbemensis (Alston)
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Isoetes hallasanensis (H.K.Choi, Ch.Kim & J.Jung)
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Isoetes hallei (Eb.Fisch. & Lobin)
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Isoetes harleyi (J.B.S.Pereira)
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Isoetes haussknechtii (Troìa & Greuter)
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Isoetes hawaiiensis (W.C.Taylor & W.H.Wagner)
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Isoetes heldreichii (Wettst.)
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Isoetes hemivelata (R.L.Small & Hickey)
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Isoetes herzogii (U.Weber)
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Isoetes heterospora (A.A.Eaton)
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Isoetes hewitsonii (Hickey)
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Isoetes hieronymi (U.Weber)
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Isoetes histrix (Bory)
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Isoetes hopei (J.R.Croft)
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Isoetes howellii (Engelm.)
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Isoetes humilior (A.Braun)
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Isoetes hyemalis (D.F.Brunt.)
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Isoetes hypsophila (Hand.-Mazz.)
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Isoetes inflata (E.R.L.Johnson)
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Isoetes jaegeri (Pitot)
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Isoetes jamaicensis (Hickey)
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Isoetes japonica (A.Braun)
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Isoetes jarmaniae (D.F.Brunt., Kantvilas & M.Garrett)
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Isoetes jeffreyi (D.M.Britton & D.F.Brunt.)
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Isoetes jejuensis (H.K.Choi, Ch.Kim & J.Jung)
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Isoetes junciformis (D.F.Brunt. & D.M.Britton)
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Isoetes karstenii (A.Braun)
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Isoetes kashyapii (M.Patel, M.N.Reddy & H.K.Goswami)
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Isoetes killipii (C.V.Morton)
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Isoetes kirkii (A.Br.)
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Isoetes labri-draconis (N.R.Crouch)
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Isoetes lacustris (L.)
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Isoetes laosiensis (C.Kim & H.K.Choi)
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Isoetes laurentiana (D.F.Brunt.)
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Isoetes lechleri (Mett.)
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Isoetes libanotica (Musselman, Bolin & R.D.Bray)
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Isoetes lithophila (N.Pfeiff.)
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Isoetes longifolia (J.B.S.Pereira)
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Isoetes longissima (Bory)
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Isoetes longpingii (Y.H.Yan, Y.F.Gu & J.P.Shu)
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Isoetes louisianensis (Thieret)
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Isoetes luetzelburgii (U.Weber in Luetzelb.)
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Isoetes malinverniana (Ces. & De Not.)
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Isoetes maritima (Underw.)
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Isoetes martii (A.Braun)
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Isoetes mattaponica (Musselman & W.C.Taylor)
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Isoetes maxima (Hickey, Macluf & Link-Pérez)
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Isoetes melanopoda (J.Gay & Durieu)
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Isoetes melanospora (Engelm.)
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Isoetes melanotheca (Alston)
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Isoetes mexicana (Underw.)
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Isoetes microvela (D.F.Brunt.)
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Isoetes minima (A.A.Eaton)
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Isoetes mississippiensis (S.W.Leonard, W.C.Taylor, Musselman & R.D.Bray)
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Isoetes mongerensis (E.R.L.Johnson)
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Isoetes montezumae (A.A.Eaton)
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Isoetes mourabaptistae (J.B.S.Pereira, P.G.Windisch, Lorscheitt. & Labiak)
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Isoetes muelleri (A.Braun)
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Isoetes naipiana (P.G.Windisch, Lorscheitt. & Nervo)
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Isoetes nana (J.B.S.Pereira)
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Isoetes neoguineensis (Baker)
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Isoetes nigritiana (A.Br.)
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Isoetes nigroreticulata (Verdc.)
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Isoetes novo-granadensis (H.P.Fuchs)
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Isoetes nuttallii (A.Braun ex Engelm.)
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Isoetes occidentalis (L.F.Hend.)
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Isoetes olympica (A.Br.)
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Isoetes orcuttii (A.A.Eaton)
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Isoetes organensis (U.Weber in Luetzelb.)
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Isoetes orientalis (Hong Liu & Q.F.Wang)
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Isoetes ovata (N.Pfeiff.)
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Isoetes pallida (Hickey)
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Isoetes palmeri (H.P.Fuchs)
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Isoetes panamensis (Maxon & C.V.Morton in Woodson & Seibert)
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Isoetes parvula (Hickey)
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Isoetes pedersenii (H.P.Fuchs ex E.I.Meza & Macluf)
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Isoetes perrieriana (Iversen)
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Isoetes philippinensis (Merr. & L.M.Perry)
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Isoetes phrygia ((Boiss.) Hausskn.)
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Isoetes pichisermollii (Eb.Fisch. & Lobin)
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Isoetes pirinica (D.F.Brunt. & D.Ivanova)
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Isoetes pitotii (Alston)
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Isoetes precocia (R.L.Small & Hickey)
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Isoetes pringlei (Underw.)
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Isoetes prototypus (D.M.Britton & Goltz)
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Isoetes pseudojaponica (M.Takamiya, Mits.Watan. & K.Ono)
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Isoetes pusilla (C.R.Marsden & Chinnock)
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Isoetes quiririensis (J.B.S.Pereira & Labiak)
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Isoetes ramboi (Herter)
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Isoetes riparia (Engelm. ex A.Braun)
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Isoetes rwandensis (Eb.Fisch. & Lobin)
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Isoetes sabatina (Troìa & Azzella)
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Isoetes saccharata (Engelm.)
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Isoetes sahyadrii (Mahab. ex L.N.Rao)
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Isoetes salishensis (D.F.Brunt.)
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Isoetes santacruzensis (J.B.S.Pereira)
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Isoetes saracochensis (Hickey)
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Isoetes savatieri (Franch.)
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Isoetes schweinfurthii (A.Br.)
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Isoetes sehnemii (H.P.Fuchs in Reitz)
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Isoetes septentrionalis (D.F.Brunt.)
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Isoetes serracarajensis (J.B.S.Pereira, Salino & Stützel)
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Isoetes shangrilaensis (Xiang Li, Yuqian Huang, X.Dai & Xing Liu)
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Isoetes sinensis (T.C.Palmer)
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Isoetes smithii (H.P.Fuchs in Reitz)
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Isoetes spannagelii (H.P.Fuchs in Reitz)
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Isoetes spinulospora (C.Jermy & Schelpe)
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Isoetes stellenbossiensis (A.V.Duthie)
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Isoetes stephanseniae (A.V.Duthie)
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Isoetes stevensii (J.R.Croft)
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Isoetes storkii (T.C.Palmer)
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Isoetes taiwanensis (De Vol)
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Isoetes tamaulipana (Mora-Olivo, A.Mend. & Mart.-Aval.)
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Isoetes tegetiformans (Rury)
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Isoetes tenella (Léman ex Desv.)
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Isoetes tennesseensis (Luebke & Budke)
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Isoetes tenuifolia (Jermy)
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Isoetes texana (Singhurst, Rushing & W.C.Holmes)
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Isoetes tiguliana (Gennari)
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Isoetes todaroana (Troìa & Raimondo)
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Isoetes toximontana (Musselman & J.P.Roux)
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Isoetes transvaalensis (C.Jermy & Schelpe)
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Isoetes triangula (U.Weber)
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Isoetes tripus (A.Braun)
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Isoetes truncata (Clute)
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Isoetes tuckermanii (A.Braun ex Engelm.)
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Isoetes tuerckheimii (Brause)
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Isoetes udupiensis (P.K.Shukla, G.K.Srivast., S.K.Shukla & P.K.Rajagopal)
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Isoetes ulei (U.Weber)
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Isoetes valida (Clute)
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Isoetes vanensis (M.Keskin & G.Zare)
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Isoetes vermiculata (Hickey)
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Isoetes virginica (N.Pfeiff.)
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Isoetes viridimontana (M.A.Rosenthal & W.C.Taylor)
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Isoetes weberi (Herter ex U.Weber)
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Isoetes welwitschii (A.Br. ex Kuhn)
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Isoetes wormaldii (Sim)
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Isoetes xiangfei (Y.H.Yan, Y.F.Gu & J.P.Shu)
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Isoetes yuhangensis (Yu C.Chen & Xing Liu)
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Isoetes yunguiensis (Q.F.Wang & W.C.Taylor)